Boyer diss 2009 1046..
Boyer diss 2009 1046.. Boyer diss 2009 1046..
southern Colorado. This locality is placed in the Tiffanian (Ti) 4 biozone (Gingerich, 1976; Lofgren et al., 2004). Simpson (1935) was the first to thoroughly describe the bones from the Mason Pocket locality attributed by him to “P. gidleyi.” Specifically, he indicated that AMNH 17379 included an atlas, other fragmentary cervical vertebrae, two thoracic vertebrae, six lumbar vertebrae, ribs, the sacrum, and two anterior caudal vertebrae. Some of these bones were illustrated (Simpson, 1935: p. 13, fig. 6). He described and illustrated the left scapula, right humerus, ulna, and radius (p. 14, fig. 7), a metacarpal and proximal phalanx (p. 15, fig. 8), right proximal femur, left calcaneum, right astragalus, and left tibia (p. 19, fig. 11). He attributed another specimen, AMNH 17409, from this locality to “P. gidleyi” as well. This specimen includes a right innominate (p. 17, fig. 9) and a left distal femur (p. 19, fig. 10). He also described the existence of many isolated intermediate phalanges, although he was not convinced that these belonged to N. gidleyi and did not figure them. Based on the morphology preserved in AMNH 17379 and 17409, Simpson concluded that N. gidleyi was likely closer to “lemurids” than “tupaiids” but that N. gidleyi must have been derived for an ecological niche very different from that of lemurid, or notharctine, euprimates. All of the bones described by Simpson and many more associated with the skeleton were observed and measured during the course of this study. Russell (1964) made a major contribution to knowledge of plesiadapid postcranial anatomy by publishing the results of his efforts at Mouras (Berru) Quarry, near the village of Cernay-Les-Reims. He provided a long list of elements recognized by him as pertaining to P. tricuspidens (Russell, 1964: p. 289-293). Some of these were indicated as being associated with single individuals. He did not, however, provide descriptions or 256
illustrations of this material, except for the claws. He compared the claws to those of the flying lemur, Cynocephalus, noting that they were similar in being mediolaterally narrow and dorsoventrally deep. The task of describing the rest of the material was left to later researchers. Simons (1964: p. 56, fig. 3) was the first to figure and discuss some of this new material. It is difficult to assess which particular specimens the illustrations in Simons (1964) were based on, because they have been reconstructed to varying degrees. However, some of the lower limb bones appear to correspond to MNHN R 408 (a complete femur) and MNHN R 410 (a fragmentary tibia), considered by Russell (1964) to be components of the same individual. It is important to note that MNHN R 410 lacks both its proximal and distal ends. Simons considered Plesiadapis to have been a treeshrew or tree squirrel-like arborealist, and reiterated Russell’s observation regarding similarity between the claws of Plesiadapis and Cynocephalus. Russell (1967) studied a slab-preserved specimen of Piton’s (1940) Menatotherium insigne from the Menat Basin in central France, and recognized its dentition as pertaining to Plesiadapis insignis. This unnumbered specimen and another lacking a skull, housed at the MNHN, allowed future researchers to estimate limb lengths and indices for P. insignis (see below). The small size of P. insignis, its similarity to early-occurring North American forms of Plesiadapis, and contextual information from the deposits that yielded the specimen, suggested to Gingerich (1976) and others that this specimen was probably contemporaneous with Tiffanian 1 index taxon Plesiadapis praecursor, making it the oldest known postcranial remains of a plesiadapid at the time (older than that from Cernay and the Mason Pocket). I was able to observe the type 257
- Page 233 and 234: DENTAL FUNCTIONAL MORPHOLOGY OF P.
- Page 235 and 236: Lower premolar molarization As indi
- Page 237 and 238: SUMMARY AND CONCLUSION The skull of
- Page 239 and 240: REFERENCES Bloch, J.I., Boyer, D.M.
- Page 241 and 242: TABLES Table 3.1. List of anatomica
- Page 243 and 244: Table 3.2. Anatomical abbreviations
- Page 245 and 246: Table 3.3. Size comparison among pl
- Page 247 and 248: Table 3.4 continued. European plesi
- Page 249 and 250: Figure 3.1. Cranium of Plesiadapis
- Page 251 and 252: Figure 3.3. Right maxillary teeth (
- Page 253 and 254: Figure 3.4. Cranium of Plesiadapis
- Page 255 and 256: Figure 3.5. Cranium of Plesiadapis
- Page 257 and 258: Figure 3.6. Cranium of Plesiadapis
- Page 259 and 260: Figure 3.8. Fragment from right nuc
- Page 261 and 262: Figure 3.9. Right promontorium of P
- Page 263 and 264: Figure 3.10. Cranium of Plesiadapis
- Page 265 and 266: Figure 3.12. Right dentary of Plesi
- Page 267 and 268: Figure 3.14. A, Plot of relief inde
- Page 269 and 270: CHAPTER 4: THE FIRST KNOWN SKELETON
- Page 271 and 272: among plesiadapiforms (e.g., Szalay
- Page 273 and 274: Institutional and collections abbre
- Page 275 and 276: CaL - capitulum (of humerus) antero
- Page 277 and 278: HSV - head shape variable = ln(DEW/
- Page 279 and 280: MSD - mid-shaft dorsoventral or ant
- Page 281 and 282: Ry - ray (as in “digit ray”) S-
- Page 283: History of descriptive study of the
- Page 287 and 288: astragalus and calcaneum was highly
- Page 289 and 290: discussion of the femur indicates t
- Page 291 and 292: supinator crests. He also noted tha
- Page 293 and 294: that it may not even be an archonta
- Page 295 and 296: unstudied material. Specifically, h
- Page 297 and 298: 5321), some metapodials (MNHN R 529
- Page 299 and 300: Gingerich and Gunnell (1992) publis
- Page 301 and 302: prehensility they provide, is an in
- Page 303 and 304: euarchontans (Fig. 1.1). Their anal
- Page 305 and 306: for comparison. These include isola
- Page 307 and 308: plesiadapid samples have the same m
- Page 309 and 310: Organization of results Each bone i
- Page 311 and 312: Bloch and Boyer (2002) and N. inter
- Page 313 and 314: clavicle reflects some basic aspect
- Page 315 and 316: Humerus Description.—The right an
- Page 317 and 318: epicondyle actually projects somewh
- Page 319 and 320: cookei is absolutely longer than an
- Page 321 and 322: tuberosity. This crest probably del
- Page 323 and 324: olecranon process to estimate its t
- Page 325 and 326: distinct, convex distal radial face
- Page 327 and 328: of the midcarpal joint), and its pr
- Page 329 and 330: (there is no evidence for more than
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illustrations of this material, except for the claws. He compared the claws to those of the<br />
flying lemur, Cynocephalus, noting that they were similar in being mediolaterally narrow<br />
and dorsoventrally deep. The task of describing the rest of the material was left to later<br />
researchers. Simons (1964: p. 56, fig. 3) was the first to figure and discuss some of this<br />
new material. It is difficult to assess which particular specimens the illustrations in<br />
Simons (1964) were based on, because they have been reconstructed to varying degrees.<br />
However, some of the lower limb bones appear to correspond to MNHN R 408 (a<br />
complete femur) and MNHN R 410 (a fragmentary tibia), considered by Russell (1964)<br />
to be components of the same individual. It is important to note that MNHN R 410 lacks<br />
both its proximal and distal ends. Simons considered Plesiadapis to have been a<br />
treeshrew or tree squirrel-like arborealist, and reiterated Russell’s observation regarding<br />
similarity between the claws of Plesiadapis and Cynocephalus.<br />
Russell (1967) studied a slab-preserved specimen of Piton’s (1940)<br />
Menatotherium insigne from the Menat Basin in central France, and recognized its<br />
dentition as pertaining to Plesiadapis insignis. This unnumbered specimen and another<br />
lacking a skull, housed at the MNHN, allowed future researchers to estimate limb lengths<br />
and indices for P. insignis (see below). The small size of P. insignis, its similarity to<br />
early-occurring North American forms of Plesiadapis, and contextual information from<br />
the deposits that yielded the specimen, suggested to Gingerich (1976) and others that this<br />
specimen was probably contemporaneous with Tiffanian 1 index taxon Plesiadapis<br />
praecursor, making it the oldest known postcranial remains of a plesiadapid at the time<br />
(older than that from Cernay and the Mason Pocket). I was able to observe the type<br />
257