Boyer diss 2009 1046..

Boyer diss 2009 1046.. Boyer diss 2009 1046..

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southern Colorado. This locality is placed in the Tiffanian (Ti) 4 biozone (Gingerich, 1976; Lofgren et al., 2004). Simpson (1935) was the first to thoroughly describe the bones from the Mason Pocket locality attributed by him to “P. gidleyi.” Specifically, he indicated that AMNH 17379 included an atlas, other fragmentary cervical vertebrae, two thoracic vertebrae, six lumbar vertebrae, ribs, the sacrum, and two anterior caudal vertebrae. Some of these bones were illustrated (Simpson, 1935: p. 13, fig. 6). He described and illustrated the left scapula, right humerus, ulna, and radius (p. 14, fig. 7), a metacarpal and proximal phalanx (p. 15, fig. 8), right proximal femur, left calcaneum, right astragalus, and left tibia (p. 19, fig. 11). He attributed another specimen, AMNH 17409, from this locality to “P. gidleyi” as well. This specimen includes a right innominate (p. 17, fig. 9) and a left distal femur (p. 19, fig. 10). He also described the existence of many isolated intermediate phalanges, although he was not convinced that these belonged to N. gidleyi and did not figure them. Based on the morphology preserved in AMNH 17379 and 17409, Simpson concluded that N. gidleyi was likely closer to “lemurids” than “tupaiids” but that N. gidleyi must have been derived for an ecological niche very different from that of lemurid, or notharctine, euprimates. All of the bones described by Simpson and many more associated with the skeleton were observed and measured during the course of this study. Russell (1964) made a major contribution to knowledge of plesiadapid postcranial anatomy by publishing the results of his efforts at Mouras (Berru) Quarry, near the village of Cernay-Les-Reims. He provided a long list of elements recognized by him as pertaining to P. tricuspidens (Russell, 1964: p. 289-293). Some of these were indicated as being associated with single individuals. He did not, however, provide descriptions or 256

illustrations of this material, except for the claws. He compared the claws to those of the flying lemur, Cynocephalus, noting that they were similar in being mediolaterally narrow and dorsoventrally deep. The task of describing the rest of the material was left to later researchers. Simons (1964: p. 56, fig. 3) was the first to figure and discuss some of this new material. It is difficult to assess which particular specimens the illustrations in Simons (1964) were based on, because they have been reconstructed to varying degrees. However, some of the lower limb bones appear to correspond to MNHN R 408 (a complete femur) and MNHN R 410 (a fragmentary tibia), considered by Russell (1964) to be components of the same individual. It is important to note that MNHN R 410 lacks both its proximal and distal ends. Simons considered Plesiadapis to have been a treeshrew or tree squirrel-like arborealist, and reiterated Russell’s observation regarding similarity between the claws of Plesiadapis and Cynocephalus. Russell (1967) studied a slab-preserved specimen of Piton’s (1940) Menatotherium insigne from the Menat Basin in central France, and recognized its dentition as pertaining to Plesiadapis insignis. This unnumbered specimen and another lacking a skull, housed at the MNHN, allowed future researchers to estimate limb lengths and indices for P. insignis (see below). The small size of P. insignis, its similarity to early-occurring North American forms of Plesiadapis, and contextual information from the deposits that yielded the specimen, suggested to Gingerich (1976) and others that this specimen was probably contemporaneous with Tiffanian 1 index taxon Plesiadapis praecursor, making it the oldest known postcranial remains of a plesiadapid at the time (older than that from Cernay and the Mason Pocket). I was able to observe the type 257

illustrations of this material, except for the claws. He compared the claws to those of the<br />

flying lemur, Cynocephalus, noting that they were similar in being mediolaterally narrow<br />

and dorsoventrally deep. The task of describing the rest of the material was left to later<br />

researchers. Simons (1964: p. 56, fig. 3) was the first to figure and discuss some of this<br />

new material. It is difficult to assess which particular specimens the illustrations in<br />

Simons (1964) were based on, because they have been reconstructed to varying degrees.<br />

However, some of the lower limb bones appear to correspond to MNHN R 408 (a<br />

complete femur) and MNHN R 410 (a fragmentary tibia), considered by Russell (1964)<br />

to be components of the same individual. It is important to note that MNHN R 410 lacks<br />

both its proximal and distal ends. Simons considered Plesiadapis to have been a<br />

treeshrew or tree squirrel-like arborealist, and reiterated Russell’s observation regarding<br />

similarity between the claws of Plesiadapis and Cynocephalus.<br />

Russell (1967) studied a slab-preserved specimen of Piton’s (1940)<br />

Menatotherium insigne from the Menat Basin in central France, and recognized its<br />

dentition as pertaining to Plesiadapis insignis. This unnumbered specimen and another<br />

lacking a skull, housed at the MNHN, allowed future researchers to estimate limb lengths<br />

and indices for P. insignis (see below). The small size of P. insignis, its similarity to<br />

early-occurring North American forms of Plesiadapis, and contextual information from<br />

the deposits that yielded the specimen, suggested to Gingerich (1976) and others that this<br />

specimen was probably contemporaneous with Tiffanian 1 index taxon Plesiadapis<br />

praecursor, making it the oldest known postcranial remains of a plesiadapid at the time<br />

(older than that from Cernay and the Mason Pocket). I was able to observe the type<br />

257

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