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Boyer diss 2009 1046..

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morphology, as well as the presence of g2 grooves, demonstrates an intratympanic route<br />

for the internal carotid plexus of P. cookei.<br />

Grooves on the promontorium<br />

Compared to other plesiadapids the g2 grooves diverge from the g1 groove<br />

unusually far proximally (Fig. 3.9): they are already distinct on the posterior septum,<br />

rather than forming farther cranially, on the promontorium. The divergence of the two g2<br />

grooves from one another on the promontorium is also unusual, and is not observed in<br />

any other plesiadapid specimens described in the Chapter 2. It is unknown whether this<br />

unusual morphology is a characteristic of P. cookei or whether it represents intraspecific<br />

variability in the course of nerves in this area (and therefore represents a pattern that<br />

could just as easily appear in an individual of another species such as Pronothodectes<br />

gaoi).<br />

Composition of bulla: sutural evidence on medial tympanic process<br />

The medial tympanic process of P. cookei is unique among the comparative<br />

sample of plesiadapids in exhibiting a “suture-like” line of foramina (Fig. 3.9D).<br />

Comparative rodent specimens of Marmota and Lagostomus (Chapter 2) show<br />

neurovasculature invading the middle ear cavity at the petrosal/ectotympanic boundary,<br />

creating a line of foramina at the sutural boundary. This extant morphological pattern<br />

makes it tempting to interpret the line of foramina in P. cookei as the remnant of a suture<br />

as well. This may or may not be correct: HRxCT data does not provide any more<br />

insights, as the boundary between the pars cochlearis of the petrosal and its medial<br />

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