Boyer diss 2009 1046..

Boyer diss 2009 1046.. Boyer diss 2009 1046..

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(Figs. 3.1, 4-6: 13), the zygomatic is 7.9 mm deep. There is no evidence of a postorbital process in this vicinity. The ventral surface of the anterior end of the zygomatic is expanded transversely for attachment of the superficial masseter (Fig. 3.5: 14) (as is the zygomatic process of the maxilla at its suture with the zygomatic). This expansion measures 2.38 mm. The total anteroposterior length of the element along its ventral margin is 24.9 mm. The posterior end has a fragment of the zygomatic process of the squamosal still attached to it. The suture between these two elements measures at least 14.2 mm. Frontal.— The frontals are visible on the dorsum of the skull; in this region the frontals clearly contact the maxillae and lacrimals (Fig. 3.1: 4, 6). The contact between the premaxillae (if any) and nasals are obscured by breakage. Furthermore, breakage makes the existence and/or nature of contacts with the palatine, orbitosphenoid, alisphenoid, and parietal difficult to assess. A metopic suture is prominent (Fig. 3.1, 6: 15) and extends from the anterior end of the frontals to the parietals. Noting some breakage in the region of nasal contact, the anteroposterior length of the frontal is 21 mm and the unilateral width, from the metopic suture to the lacrimal suture, is roughly 15.3 mm. The frontals exhibit distinct ridges (temporal crests) that run medially from the anterolateral part of the bones, where they contact the lacrimals, toward the metopic suture. The temporal crests meet the metopic suture at the posterior end of the frontals, where they contact the parietals (Fig. 3.1: 16). Thus, they would have formed a distinct “trigon” on the forehead. There is no sign of postorbital processes on these bones. HRxCT data reveal the frontal as a thin plate of bone anteriorly and show that the bone thickens posteriorly and is densely trabeculated. No major diploic cavities are 190

identifiable. No ethmoid foramina could be identified. At the parietal’s most anterior point of contact with the frontal, it reaches a few millimeters beyond the apex of the converging temporal crests, to the point of maximum mediolateral constriction of the orbitotemporal region (Fig. 3.1, 4: 17). This suture is difficult to see with the naked eye or light microscopy; however, HRxCT imagery reveals its position and form more distinctly. The parietal onlapped the frontal (Fig. 3.7: 18). Thus, although much of the neurocranium has an “outer shell” of parietal, the frontal extends quite far posteriorly. Most likely, the frontal forms at least the anterior part of the endocranial surface, the topology of which reflects that of the brain, mainly the olfactory bulbs (Fig. 3.5: 19). In fact, the posterior part of the frontal has its endocranial surface and the endocranial expression of its suture with the parietal exposed (Fig. 3.5: 20). This region is described by Gingerich and Gunnell (2005). Palatine.— The palatal processes of the palatines are visible and well preserved (Fig. 3.5). The base of the left pterygoid process is preserved (Fig. 3.5: 21). More of the posterior parts also may be present, but shifted dorsally and posteriorly into the neurocranium (Fig. 3.5). On the palate, the palatines terminate at the choanae in a swollen rim of bone, also referred to as a postpalatine torus (Fig. 3.5: 22). The outer margin of the torus reaches the lateral margin of the pterygoid processes. Furthermore, the outer margin of the torus is sharply angled, with the anterior part of it following a straight, transversely-running course, and the lateral parts following a straight, anteroposteriorly-running course. Prior to shifting of the contralateral palatines the inner margin of the horizontal process of the palatine, which forms the direct boundary to the choanae, would have been biconcave with a postpalatine spine present (Fig. 3.5: 23). This 191

identifiable. No ethmoid foramina could be identified. At the parietal’s most anterior<br />

point of contact with the frontal, it reaches a few millimeters beyond the apex of the<br />

converging temporal crests, to the point of maximum mediolateral constriction of the<br />

orbitotemporal region (Fig. 3.1, 4: 17). This suture is difficult to see with the naked eye<br />

or light microscopy; however, HRxCT imagery reveals its position and form more<br />

distinctly. The parietal onlapped the frontal (Fig. 3.7: 18). Thus, although much of the<br />

neurocranium has an “outer shell” of parietal, the frontal extends quite far posteriorly.<br />

Most likely, the frontal forms at least the anterior part of the endocranial surface, the<br />

topology of which reflects that of the brain, mainly the olfactory bulbs (Fig. 3.5: 19). In<br />

fact, the posterior part of the frontal has its endocranial surface and the endocranial<br />

expression of its suture with the parietal exposed (Fig. 3.5: 20). This region is described<br />

by Gingerich and Gunnell (2005).<br />

Palatine.— The palatal processes of the palatines are visible and well preserved<br />

(Fig. 3.5). The base of the left pterygoid process is preserved (Fig. 3.5: 21). More of the<br />

posterior parts also may be present, but shifted dorsally and posteriorly into the<br />

neurocranium (Fig. 3.5). On the palate, the palatines terminate at the choanae in a<br />

swollen rim of bone, also referred to as a postpalatine torus (Fig. 3.5: 22). The outer<br />

margin of the torus reaches the lateral margin of the pterygoid processes. Furthermore,<br />

the outer margin of the torus is sharply angled, with the anterior part of it following a<br />

straight, transversely-running course, and the lateral parts following a straight,<br />

anteroposteriorly-running course. Prior to shifting of the contralateral palatines the inner<br />

margin of the horizontal process of the palatine, which forms the direct boundary to the<br />

choanae, would have been biconcave with a postpalatine spine present (Fig. 3.5: 23). This<br />

191

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