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Boyer diss 2009 1046..

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primates, as well as treeshrews and elephant shrews, according to MacPhee (1981). It<br />

has also been identified in Ignacius graybullianus (Bloch and Silcox, 2001) (Fig. 2.38).<br />

Either the s2 or s3 is equivalent to the medial secondary septum of MacPhee<br />

(1981), while the other one cannot be analogized with morphology present in the sample<br />

of primates and treeshrews studied by him. The hypothesis that the s2 is more nearly<br />

equivalent to the medial secondary septum is favored here as discussed below.<br />

The g1 groove is clearly related to the internal carotid plexus. This is supported<br />

by its anatomical relationships and its typical morphology. Specifically, it is located on<br />

the lateral aspect of the posterior septum and ventral apex of the pars cochlearis of the<br />

petrosal. It thus leads directly anterior from the internal carotid canal formed in the floor<br />

of the posterior septum. Furthermore, the fact that it often appears as a pair of grooves is<br />

consistent with a relationship to the internal carotid plexus, because the internal carotid<br />

plexus commonly exists as two major nerve bundles (e.g., MacPhee, 1981; Conroy and<br />

Wible, 1978; Wible, 1993).<br />

The g2 groove on the petrosal, which is usually present, is also interpreted as<br />

having held contents of the internal carotid plexus. This groove begins from where g1<br />

reaches the promontorium, and always approaches the lateral side of the anterior septum.<br />

MacPhee (1981) and other authors (e.g., Conroy and Wible, 1978) demonstrate that the<br />

internal carotid plexus follows the anterior septum (and thus leads towards the tubal<br />

canal) in various lemuroid euprimates, treeshrews and macroscelidean insectivores. My<br />

own observations reveal that the internal carotid plexus of Capra also leads towards the<br />

tubal canal. These anatomical associations and the fact that g2 is sometimes present as a<br />

pair of parallel grooves strongly suggest that it relates to the internal carotid plexus.<br />

77

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