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Flora of Dominica, Part 2 - Smithsonian Institution Libraries

Flora of Dominica, Part 2 - Smithsonian Institution Libraries

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66 CWSIACEAE SMITHSONIAN CONTRIBUTIONS TO BOTANY<br />

2. Leaves lanceolate; petiole without a pit at the base;<br />

flowers in elongate axillary racemes; sap clear. ....<br />

.......................... Mariih<br />

2. Leaves ovate; petiole with margined pit at base; flowers<br />

in fascicles or short cymes; sap yellow.<br />

3. Flowers in lateral fascicles; non-prop-rooted trees <strong>of</strong><br />

north & eastern slopes near sea. ..... Garcinia<br />

3. Flowers in terminal cymes; prop-rooted trees <strong>of</strong><br />

interior rainforests ............. Tovomita<br />

1. Venation undifferentiated, striate.<br />

4. Flowers in terminal cymes; leaves strongly obovate;<br />

petiole with a margined pit at base ....... Clusia<br />

4. Flowers axillary or in terminal fascicles; leaves elliptic<br />

to weakly obovate; petiole without margined pit at base.<br />

5. Leaves acute; flowers red, in terminal fascicles; trees<br />

with prop roots ............. Symphonia<br />

5. Leaves rounded; flowers white to yellow, axillary;<br />

trees without prop roots.<br />

6. Flowers in axillary cymes; leaves with inconspicuous<br />

glandular lines ......... Calophyllum<br />

6. Flowers in axillary fascicles or solitary; leaves<br />

conspicuously glandular-punctate ... Mammea<br />

Calophyllum Linnaeus<br />

Calophyllum antillanurn<br />

Calophyllwn antillanurn Britton in Brinon & Wilson,1924.5584.<br />

Calophyllwn calaba Jacquin, 1763:269, pl. 165, non Linnaeus.<br />

Calophyllwn brasiliense Cambessedes var. antillanwn (Brittm) Standley,<br />

1932:7.-Little & Wadsworth, 19M348, pl. 161.<br />

Calophyllwnjacquinii Fawcett & Rendle, 1926,5200, nom. illeg. [incl. type <strong>of</strong><br />

Calophyllwn antillanwn Britton, 19241.<br />

Calophyllwn calaba sensu aucn. non haeus.-Furtado, 1941.-Howard,<br />

1962398.--Lourteig & Fosberg, 1985.-Howard, 1989,5320.<br />

Galba.<br />

Shrub or tree to 30 m; sap white; leaves elliptic or<br />

elliptic-oblong, to 15 cm x 6 cm; petals white; drupe globoid,<br />

to 2.5 cm wide.<br />

West Indies; common in coastal thickets along north and east<br />

coast <strong>of</strong> <strong>Dominica</strong> to 30 m, probably planted elsewhere:<br />

Calibishie area (Ernst 1552,1687,1828, Hodge 3180), Clarke<br />

Hall (Nicolson 2006, Stern & Wasshausen 2442), Portsmouth<br />

(Hodge 533), Rosalie-La Plaine (Chambers 2725, Ernst 191 7,<br />

Wilbur 8165,8327), Salybia (Hodge 3199). Flowering February-July,<br />

fruiting August-April.<br />

Used as windbreak and for timber.<br />

The name Calophyllum calaba Linnaeus has mixed usage in<br />

the sense <strong>of</strong> a neotropical element and as at least one Old World<br />

element. I think Stevens (in Manilal, 1980:168-176;<br />

1980:256) was correct in concluding that the name must be<br />

typified on an Old World element, essentially as previously<br />

concluded by Britton (l.c.) and Fawcett and Rendle (1.c). I do<br />

not see that the Plumier element (source <strong>of</strong> the species epithet),<br />

composed only <strong>of</strong> flowers and fruits, can support (be used as<br />

lectotype) for a name validated by a nomen specificwn<br />

legitimum that only pertains to leaf characters (“foliis ovatis<br />

obtusis”). Some have suggested abandoning the Linnaean<br />

name as a nomen dubium because <strong>of</strong> its mixed usage. For<br />

further discussion see Nicolson et al. (1988:81).<br />

Clusia Linnaeus<br />

1. Epiphytic shrub or tree (dry west coast or rainforest);<br />

petioles 1-2 cm long, fruit and flowers l(-3), individually<br />

C. major<br />

pedicellate ....................<br />

1. Shrubby tree (mossy forest); leaves k sessile; fruit and<br />

flowers many, individually sessile ....... C. mangle<br />

Clusia major<br />

Clusia major Linnaeus, 1753:509.-Howard, 1989,5:323.<br />

Clusia alba Jacquin. 1760:34.-Grisebach, 1860:107.<br />

Clvsia plukenetii sensu Hodge, 1954, pp. 33, 40, and Hodge & Taylor,<br />

1957585, non Urban.<br />

Caquelin, fige kaklin, pomme gros, lagalie, yabrico maron.<br />

Shrub or epiphytic strangler to 20 m; sap white or yellow,<br />

sticky; petals white; staminate material rare; fruit ovoid to<br />

ellipsoid, dark purple, 3.5-6.0 cm x 5-6 cm, on a peduncle to<br />

3 cm long, possibly indehiscent.<br />

Lesser Antilles; in <strong>Dominica</strong> common in dry coastal areas to<br />

montane rain-forests, 35-800 m: Baiac (Whitefoord 3778),<br />

Clarke Hall (Nicolson 1828), Freshwater Lake area (Chambers<br />

2682, Ernst 1779, Smith 10218, Wilbur 7442), Grand Savanne<br />

(Stern& Wasshausen 2460), La Chaudiere (Hodge 3668), Pont<br />

Casd area (Ernst 1813, 1859, Hodge 1205, Wilbur 7846),<br />

Salybia (Hodge 3206, 3251), Sylvania (Cooper 63, Hodge<br />

535). Syndicate (Whitefoord 4386). Flowering and hiting<br />

appear to be continuous but Howard (1.c.) reports only one male<br />

specimen and no dehiscent fruits have been seen in the field or<br />

herbarium.<br />

Aerial roots used for basket making and the sap for bird lime<br />

(Hodge and Taylor, 1957:585).<br />

Clusia major Linnaeus was extremely heterogeneous, as<br />

published, but has been lectotypified (Howard, 1962392, fig.<br />

1) on Clusiaflore albo, fructu coccineo Plumier (1703:21, pl.<br />

87: fig. 1). This same plate was cited in the publications <strong>of</strong><br />

Clusia alba Jacquin (1760 and 1763). Linnaeus (1763:1495)<br />

adopted Jacquin’s binomial and dropped his own earlier<br />

binomial, an illegitimate procedure under the present Code.<br />

Planchon and Triana misidentified Imray 358 (K) from<br />

<strong>Dominica</strong> as Clusia minor Linnaeus, a species <strong>of</strong> Central and<br />

northern South America and the Greater Antilles. Planchon and<br />

Triana’s misidentification is the source <strong>of</strong> erroneous subsequent<br />

citations <strong>of</strong> C. minor for <strong>Dominica</strong>.<br />

Clusia plukenetii Urban (1908, 5:432) <strong>of</strong> Martinique, St.<br />

Lucia and Barbados has a globoid fruit borne on a peduncle to<br />

13 cm long.

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