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Flora of Dominica, Part 2 - Smithsonian Institution Libraries

Flora of Dominica, Part 2 - Smithsonian Institution Libraries

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NUMBER 77 BIGNONIACBAE 51<br />

The binomial Tecomu leucoxylon Martius ex A.P. Candolle<br />

(1845, 9:219) is a legitimate name under Article 72 (Note)<br />

(ZCBN), with priority dating from 1845, not 1763. De Candolle<br />

and later authors (Grisebach, 1862447; Urban, 1911, 4565)<br />

also included elements <strong>of</strong> what would not only appear to be T.<br />

heterophyllu (pan-Antillean) but also T roseu (South America)<br />

and T ripuriu (Jamaica).<br />

Bignoniu pentuphyllu Linnaeus (1763:870) comes next.<br />

Sandwith (1953:453) argued that this constitutes an illegitimate<br />

renaming <strong>of</strong> Bignoniu leucoxylon Linnaeus (1753:624) and<br />

thus falls into the synonymy <strong>of</strong> Tubebuiu ripuriu (Jamaican<br />

endemic). The binomial Tecomu pentuphyllu Jussieu ex A.P.<br />

Candolle (1845, 9:217) is a legitimate name under Article 72<br />

(Note) (ZCBN) with priority from 1845, not 1763. It is possible<br />

that a lectotype <strong>of</strong> Tecomu pentuphyllu could be selected that<br />

could be identified with the pan-Antillean species (I: heterophyllu),<br />

but de Candolle apparently also included the Lesser<br />

Antillean endemic (I: pullidu). In any case, Grisebach<br />

(1861:447) and Urban (1911, 4565) indiscriminately used<br />

Tecomu pentuphyllu for elements <strong>of</strong> all four species.<br />

Ruputiu (Rutaceae!) heterophyllu A.P. Candolle<br />

(182k 153) is based on sterile material from Puerto Rico. The<br />

type (micr<strong>of</strong>iche) corresponds to our concept <strong>of</strong> the pan-<br />

Antillean species, for which we accept the name Tubebuiu<br />

heterophyllu (A.P. Candolle) Britton (191548).<br />

Finally, Bignoniu pullidu Lindley (1826b) is based on<br />

unifoliolate material cultivated in St. Vincent. This is what we<br />

regard as the Lesser Antillean endemic, for which we accept the<br />

name Tubebuiu pullidu (Lindley) Miers. This name has been<br />

generally used for other taxa, particularly the pan-Antillean<br />

taxon (Britton and Wilson, 1925, 6:196, Gooding et al.,<br />

1965389).<br />

We reemphasize that our acceptance <strong>of</strong> the species rank for<br />

the two <strong>Dominica</strong>n taxa is provisional. On <strong>Dominica</strong> they are<br />

very distinct in aspect and habitat but material from other<br />

islands is not so easily divided. Stehl6 (1945338; 1946a:32)<br />

recognized three subspecies within a variable concept <strong>of</strong> I:<br />

heterophyllu. Little and Wadsworth (1964:498) went further<br />

and accepted I: heterophyllu as a variable species without<br />

subspecies. A revision <strong>of</strong> Antillean Tabebuieae, particularly<br />

accounting for the Cuban taxa, is needed.<br />

1. Leaflets (3-)5, west (leeward) coast . . . I: heterophylla<br />

1. Leaflets 1(-3); east (windward) coast . . . . . I: pallida<br />

Tabebuia heterophylla<br />

Tabebuia heterophylla (A.P. Candolle) Britton. 1915:18.-Little & Wadswod,<br />

1964:198, pl. 236.<br />

Rap& heterophylla A.P. Candolle, 1822~153; 1824, 1:734.<br />

Tecoma pentaphylla Jussieu ex A.P. Candolle, 1845, 9:217.-Urban, 1911,<br />

4565.<br />

Tabebuia pentaphylla (A.P. Candolle) Hemsley, 1882, 2:495.-Urban, 1921,<br />

8:641.<br />

Tabebuia pallida subsp. pentaphylla (A.P. Candolle) Stehlb, 1945:329,338.<br />

Tabebuia heterophylla subsp. pallida sensu Stehlb. 1946a:32, as to description,<br />

not as to the Lindley type.<br />

White cedar, poirier blanc.<br />

Tree to 6(-20) m; leaves (3-)5-foliolate, leaflets to 9(-12)<br />

cm x 5.5 cm, apex obtuse to emarginate and base rounded,<br />

petioles to 5 cm, petiolules to 3 cm; calyx 1.5 cm long,<br />

irregularly lobed; corolla to 8 cm long, pale violet with<br />

yellowish center, lobes flattened, to 2 cm long; fruits terete, to<br />

20 cm x 1 cm, apex attenuate; seeds flattened, 0.6 cm x 1 cm,<br />

hyaline wings to 2 cm long.<br />

Antillean (except Jamaica); common in dry scrub along west<br />

coast <strong>of</strong> <strong>Dominica</strong>: Cabrit (Smith 10310), Grand Savanne<br />

(Beard 246), Hungry Hill (Whitefoord 4483,Loubiere (Hodge<br />

3864), Mero (Stern & Wusshausen 2437), Morne Bruce<br />

(Fuirchild 283I), Morne Daniel (Webster 13291), Pointe<br />

Ronde (Hodge 2660,2755), Salisbury (Chambers 2801), Scotts<br />

Head (Hodge 1603, Kimber 884, Wilbur 7589), Tarou Cliffs<br />

(Nicolson 1858). Flowering January-June, fruiting in July.<br />

Tabebuia pallida<br />

Tabebuia pallida (Lindley) Miers, 1863:199.<br />

Bignonia pallida Lindley, 1826b.<br />

Tabebuia Mnicensis Urban, 1924:308.<br />

Tabebuia pallida subsp. dominicensis (Urban) Stehlb,1945:330,338.<br />

Tabcbuia heterophylla subsp. dm’nicensis (Urban) Stehlb,196:33.<br />

Poirier.<br />

Tree to lo(-30) m; leaves usually simple with apex rounded<br />

to obtuse, to 20 cm x 9 cm, petioles to 5 cm, or when rarely<br />

3-fOliOhte, leaflets to 12 cm x 5.5 cm, petioles to 9 cm and<br />

petiolules to 3 cm; pedicels 3 cm, slender; flowers and fruits as<br />

above; seeds as above but perhaps k larger.<br />

Guadeloupe, Martinique, St. Vincent, Barbados, Grenadines,<br />

and St. Lucia; common in <strong>Dominica</strong> in the eastern (windward)<br />

coastal forests: Au Park (Red 20270), Bout Sable Bay<br />

(Nicolson 1983, L’Anse Noire (Wilbur 7518), Hatton Garden<br />

(Hodge 3055), La Plaine (Rumuge s.n., type at K), Marigot<br />

(Hodge 839), Petit Soufriere Bay (Stern & Wusshuusen 2477),<br />

St. Davids Bay (Emst 1470, Wilbur 7987), St. Saveur<br />

(Chambers 2549), Salybia (Stehle 6091, 6387). Flowering<br />

January-May; fruit formed in many months, apparently mostly<br />

in July.<br />

Caribs use the wood <strong>of</strong> this (perhaps also the other species)<br />

for ship building and other purposes (Hodge and Taylor,<br />

1957608). Adjanohoun et al. (198557, pl. 23) reported<br />

medicinal usage <strong>of</strong> what probably is a composite <strong>of</strong> the two<br />

taxa recognized here.<br />

Tanaecium Swartz<br />

Tanaecium crucigerum<br />

TaMeciwn crucigerwn B. Smann, 1856:27.4enuy, 1975: 122.<br />

Scrubby vine to 3 m; leaflets elliptic, apex rounded or

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