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Flora of Dominica, Part 2 - Smithsonian Institution Libraries

Flora of Dominica, Part 2 - Smithsonian Institution Libraries

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30 ASTERACEAE SMITHSOMAN CONTRIBUTIONS TO BOTANY<br />

I am grateful to Dr. H.E. Robinson who helped with the key<br />

and in other ways.<br />

Cultivated Genera <strong>of</strong> Asteraceae<br />

Artemisia absinthium Linnaeus and Artemisia vulgaris<br />

Linnaeus were cited as cultivated in the Antilles by Adjanohoun<br />

et al. (1985:71, pl. 37-38), suggesting both are on<br />

<strong>Dominica</strong> (new records, if true).<br />

Ayapana triplinervis (Vahl) King & Robinson was attributed<br />

to <strong>Dominica</strong> as Eupatorium triplinerve Vahl by Vdez<br />

(1957:8 l), extrapolated from Britton and Wilson’s citation<br />

(1925, 6:289), “Martinique; Guadeloupe.” It has been cultivated<br />

on the French Islands and escaped but no material from<br />

<strong>Dominica</strong> has been seen. It has opposite, entire, lanceolate<br />

leaves tapering to a subsessile base. However, Adjanohoun et<br />

al. (198581, pl. 48) reported it as widely cultivated for its<br />

medicinal usages, another new record for <strong>Dominica</strong> if there.<br />

Cosmos caudatus Kunth, with pink ligules, was reported by<br />

Domin (1930d79) as collected by Imray.<br />

Cosmos sulphureus Cavanilles, with yellow or orange<br />

ligules, was reported by Adjanohoun et al. (198575, pl. 42).<br />

These have opposite, pinnatisect leaves and beaked achenes<br />

with barbed awns.<br />

Helianthus annuus Linnaeus, the sunflower, may be cultivated<br />

on <strong>Dominica</strong> but no specimens have been seen. It has<br />

alternate, dentate and rough leaves and a large head 30 or more<br />

cm across.<br />

Launaea intybacea (Jacquin) Beauverd is reported as<br />

“occasionally met in the Lesser Antilles” by Adjanohoun et al.<br />

(198585, pl. 51). This, if true for <strong>Dominica</strong>, would be a new<br />

record for the island.<br />

Tugeres erecta Linnaeus (African marigold) was reported as<br />

collected by Domin (193N79). Tagetes patula Linnaeus<br />

(French marigold) was illustrated by Adjanohoun et al.<br />

(198591, pl. 58). These species have opposite or alternate,<br />

pinnatisect leaves and connate involucral bracts. Howard<br />

(1989,6601) treated these as synonyms.<br />

Zinnia elegans Jacquin was collected in cultivation by<br />

Domin (193N73). This and other cultivated zinnias have<br />

opposite, sessile, and entire leaves.<br />

Excluded Genera <strong>of</strong> Asteraceae<br />

Borrichia arborescens (Linnaeus) A.P. Candolle was ataibuted<br />

to <strong>Dominica</strong> by Velez (1957:80). No collections have been<br />

seen and it is unlikely that this shrubby coastal calciphile (with<br />

opposite, entire, oblanceolate leaves) survives on <strong>Dominica</strong>.<br />

Helenium quadridentatum Labillardiere was credited to<br />

<strong>Dominica</strong> by Adjanohoun et al. (1985:83, pl. 49) but no<br />

material has been seen.<br />

Lagascea mollis Cavanilles was attributed to <strong>Dominica</strong> as<br />

Nocca mollis (Cavanilles) Jacquin by Velez (1957:80),<br />

extrapolated from Britton and Wilson’s citation (1925,6:300),<br />

“Anguilla to Barbados.” Although it is weedy, the few<br />

collections made on the French islands indicate that it is<br />

introduced and rare there. No collections from <strong>Dominica</strong> have<br />

been seen. Its leaves are generally opposite. Like Rolandra it<br />

has 1-flowered heads gathered in head-like glomerules but,<br />

unlike Rolandra, with a pseudo-involucre. The true involucral<br />

bracts are partly united.<br />

1. Leaves opposite.<br />

2. Pappus bristles with prominent retrorse barbs .............. Bidens<br />

2. Pappus without retrorse barbs.<br />

3. Leaves compound or deeply pinnatifid ............... Ambrosia<br />

3. Leaves simple.<br />

4. Fruits spiny, at least one large terminal spine hooked ............<br />

.............................. Acanthospermum<br />

4. Fruits not spiny.<br />

5. Leaf margins, near base, pectinate (with elongate bristles); lower leaf<br />

surface with large, dark glandular “dots” ............. Pectis<br />

5. Leaf margins not pectinate; lower surface without glandular “dots” or, if<br />

any, then inconspicuous.<br />

6. Heads with receptacular bracts (pales, sometimes bristles); heads<br />

heterogamous (outer florets female or neuter), corollas <strong>of</strong>ten yellow.<br />

7. Heads conspicuously radiate (ray-florets longer than involucre).<br />

8. Pappus <strong>of</strong> many plumose, capillary bristles ........ Tridux<br />

8. Pappus lacking or a cup <strong>of</strong> united scales.<br />

9. Involucral bracts erect, longer than achenes and receptacular<br />

pales.. ....................... Wedelia<br />

9. Involucral bracts deflexing, shorter than achenes and receptacular<br />

pales. ....................... Wulffia<br />

7. Heads inconspicuously radiate (ray-florets shorter than involucre).

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