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18 SOCIETY OF VERTEBRATE PALEONTOLOGY, MEMOIR 3<br />

X can nasolac<br />

I\<br />

pneu fen suborb<br />

nas<br />

ap caud<br />

can<br />

1 cm<br />

proc max 4," fac a i max -<br />

FIGURE 12. Plateosaurus engelhardti, facial skeleton. A, reconstruction<br />

<strong>of</strong> snout in left lateral view. B, same in ventral view. C, right nasal<br />

<strong>of</strong> AMNH 6810 in ventromedial view. D, stereophotographs and interpretive<br />

drawing <strong>of</strong> right maxilla, lacrimal, and palatine <strong>of</strong> AMNH 6810<br />

in dorsal and somewhat medial view. The nasal recess shown in C caps<br />

the hiatus in D. A,B modified after Galton (1990).<br />

I<br />

i~ In can<br />

jUY neurovas int<br />

FIGURE 11. Iguanodon atherjieldensis, facial skeleton. A, Snout in<br />

left lateral view (modified from Noman,1986). B, left maxilla, jugal,<br />

lacrimal, and prefrontal <strong>of</strong> BMNH R5764 in medial view. Arrow shows<br />

the course <strong>of</strong> the nasolacrimal duct.<br />

tending into the premaxillae as well. These foramina and canals<br />

likely conveyed branches <strong>of</strong> the maxillary nerve and accompanying<br />

vessels. Their positions dorsal or dorsolateral to the<br />

putative muscular fossa on the palatine are consistent with the<br />

dorsal pterygoideus muscle being restricted to the floor <strong>of</strong> the<br />

antorbital cavity.<br />

Dinosauria-Among Omithodira, comparable data are abundant<br />

for dinosaurs, but less so for other taxa. Good material is<br />

available for the basal members <strong>of</strong> many <strong>of</strong> the major clades<br />

<strong>of</strong> Ornithischia (Sereno, 1986). The basal omithischian Leso-<br />

thesaurus diagnosticus (Weishampel and Witmer, 1990a; Sereno,<br />

1991a) provides information on both the existence and<br />

position <strong>of</strong> a dorsal pterygoideus and the course <strong>of</strong> the maxillary<br />

neurovasculature. The dorsal surface <strong>of</strong> the palatine<br />

(BMNH R8501, RUB17) has a well developed excavation extending<br />

rostrally up to the lateral aspect <strong>of</strong> the choana (Fig. 7C,<br />

D). This excavation is here interpreted as a muscular fossa and<br />

is bounded rostrally .by a strong ridge that extends from the<br />

maxillary contact (where it borders the choana) rostrodorsomedially<br />

to the contact with the vomer. The palatine (BMNH<br />

RUB17) has an additional deep fossa rostral to this ridge and<br />

dorsomedial to the choana and directly opposite the internal<br />

antorbital fenestra that must be associated with either the nasal<br />

or antorbital cavity. The muscular fossa is restricted to the caudoventral<br />

portion <strong>of</strong> the antorbital cavity. The proximity <strong>of</strong> the<br />

ridge bordering the fossa to the medial edge <strong>of</strong> the lacrimoprefrontal<br />

wall (BMNH R8501) suggests that the postnasal fenestra<br />

was narrow in this area and would make rostral passage <strong>of</strong> a<br />

muscular slip unlikely. The course <strong>of</strong> the maxillary neurovas-

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