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Nomenclatural changes in the family Ciidae - Acta Entomologica ...

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136<br />

JELÍNEK: <strong>Nomenclatural</strong> <strong>changes</strong> <strong>in</strong> <strong>the</strong> <strong>family</strong> <strong>Ciidae</strong><br />

Results<br />

1. Eridaulus Thomson, 1859, and Eridaulus Thomson, 1863<br />

Problem of <strong>the</strong> genus-group name Eridaulus was commented on by LOHSE (1964) and<br />

LAWRENCE (1965). LOHSE (1964) po<strong>in</strong>ted out that Cis comptus Gyllenhal, 1827, was <strong>the</strong> type<br />

species of Eridaulus and that <strong>the</strong> name Eridaulus was later mis<strong>in</strong>terpreted and used as <strong>the</strong><br />

name for <strong>the</strong> group of species related to Cis nitidus, to which Cis comptus does not belong.<br />

On <strong>the</strong> contrary, LAWRENCE (1965: 280) summarized his op<strong>in</strong>ion <strong>in</strong> <strong>the</strong> follow<strong>in</strong>g two po<strong>in</strong>ts:<br />

(i) The genus Eridaulus Thomson, 1863, orig<strong>in</strong>ally <strong>in</strong>cluded two species: Anobium nitidum<br />

Fabricius, 1792, and Cis jacquemarti Mellié, 1849. (ii) The only worker to designate a type<br />

for Eridaulus was ABEILLE DE PERRIN (1874), who selected Cis comptus Gyllenhal, 1827. This<br />

species, however, was not orig<strong>in</strong>ally <strong>in</strong>cluded <strong>in</strong> <strong>the</strong> genus, and is <strong>the</strong>refore unavailable as a<br />

type. In <strong>the</strong> same work, ABEILLE DE PERRIN (1874) designated Cis nitidus as <strong>the</strong> type species<br />

of Entypus Redtenbacher. 1847, mak<strong>in</strong>g a similar error. Later, LAWRENCE (1971) designated<br />

Anobium nitidum Fabricius, 1792 [<strong>in</strong> fact Kateretes castaneus Herbst, 1793 misidentifi ed<br />

as Anobium nitidum Fabricius, 1792, see below] as <strong>the</strong> type species of Eridaulus Thomson,<br />

1863.<br />

However, LAWRENCE (1965) overlooked <strong>the</strong> fact that Thomson himself designated <strong>the</strong> type<br />

species of his new genera <strong>in</strong> <strong>the</strong> fi rst volume of his Skand<strong>in</strong>aviens Coleoptera (THOMSON<br />

1859). Thus his new genus-group names are available by <strong>in</strong>dication (ICZN 1999, Article<br />

12.2.5) with <strong>the</strong> date 1859, even though <strong>the</strong> formal descriptions of relevant genera appeared<br />

later <strong>in</strong> subsequent volumes of <strong>the</strong> book. In <strong>the</strong> fi rst volume THOMSON (1859: 91) designated<br />

Cis comptus Gyllenhal, 1827, as <strong>the</strong> type species of Eridaulus (orig<strong>in</strong>al designation) and Cis<br />

nitidus as <strong>the</strong> type species of Entypus Redtenbacher, 1847 (subsequent designation). ABEILLE<br />

DE PERRIN (1874) and LOHSE (1964) thus only accepted <strong>the</strong> previous designations by THOMSON<br />

(1859), which rema<strong>in</strong>ed neglected by most subsequent authors. Whereas <strong>the</strong> designation of<br />

Cis nitidus as <strong>the</strong> type species of Entypus is <strong>in</strong>deed erroneous because <strong>the</strong> species was not<br />

orig<strong>in</strong>ally <strong>in</strong>cluded <strong>in</strong> Entypus by REDTENBACHER (1847), Cis comptus is perfectly valid as<br />

<strong>the</strong> type species of Eridaulus Thomson, 1859, a junior subjective synonym of Cis Latreille,<br />

1796 (type species Dermestes boleti Scopoli, 1763). Eridaulus Thomson, 1863, conta<strong>in</strong><strong>in</strong>g<br />

Cis nitidus and Cis jacquemarti, can be understood as a mis<strong>in</strong>terpretation of Eridaulus Thomson,<br />

1859, and <strong>the</strong> subsequent designation of Cis nitidus as its type species by LAWRENCE<br />

(1971) is <strong>in</strong>valid.<br />

Species related to Cis nitidus auct. (non Fabricius, 1792) (see below) and correspond<strong>in</strong>g<br />

to Eridaulus sensu THOMSON (1863) are morphologically dist<strong>in</strong>ct from o<strong>the</strong>r species of Cis<br />

Latreille, 1796 (type species Dermestes boleti Scopoli, 1763), and seem to represent a monophyletic<br />

group which was given a rank vary<strong>in</strong>g between genus and species-group by various<br />

authors. THOMSON (1863) decribed it as a dist<strong>in</strong>ct genus but it was more frequently classifi ed<br />

as a subgenus of Cis, e.g. by KIESENWETTER (1898), SCHILSKY (1900), REITTER (1901) and<br />

ABDULLAH (1973). LAWRENCE (1965) argued <strong>in</strong> favour of its generic rank but later (LAWRENCE<br />

1971) reduced it to <strong>the</strong> Cis nitidus species-group. This concept was accepted e.g. by LOHSE<br />

(1965), KAWANABE (1997) and ORLEDGE & BOOTH (2006). Never<strong>the</strong>less, LOHSE & LUCHT<br />

(1992) aga<strong>in</strong> proposed subgenus rank for it. However, when this group is treated as a genus-

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