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million cells/g soil<br />

25<br />

20<br />

15<br />

10<br />

5<br />

0<br />

Burial place “Abganerovo”, light-chestnut soil<br />

SA NA RM<br />

PS-1 PS-2 PS-3 PS-4 MS<br />

Burial place “Kolobovka”, light-chestnut soil<br />

million cells/g soil<br />

25<br />

20<br />

15<br />

10<br />

5<br />

0<br />

PS-5 PS-6 PS-7 MS<br />

SA – soil agar (microorganisms consuming nutrient elements from poor environment)<br />

NA – nitrite agar (microorganisms consuming humus substances)<br />

RM – rich medium (microorganisms consuming an easily available organic matter)<br />

Fig. 1. Quantity of microorganisms of various trophic clusters in the horizon B2 in<br />

the paleosols (PS) and modern surface soils (MS)<br />

reduction of the first and second parameters, whereas the index of oligothrophicity<br />

grows up. This index expresses the ability of microbial community to use nutrient<br />

elements from poor environment: the poorer nutrient conditions are the higher<br />

index of oligothrophicity is.<br />

The temporary and spatial variability of trophic structure of soil microbial<br />

community in several regions of the steppe zone has been described. For instance,<br />

during the last 3600 years in the saline chestnut soils in the river Esaulovsky Aksay<br />

valley the microorganisms consuming nutrient elements from poor environment<br />

prevailed (74-97%) (Table 1). These microorganisms also prevailed (69-77%) in<br />

the light-chestnut soils developed on the saline loams in the Lower Volga region<br />

under more arid climate. In the light-chestnut soils, developed under conditions of<br />

less saline soil-forming rocks and higher precipitation on the watershed areas of<br />

Ergeny Upland the microorganisms growing on the rich organic medium prevailed.<br />

44

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