Zooplankton of the open Baltic: Extended Atlas - IOW
Zooplankton of the open Baltic: Extended Atlas - IOW
Zooplankton of the open Baltic: Extended Atlas - IOW
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3. METHODS OF COLLECTING AND ANALYSING<br />
ZOOPLANKTON IN THE BALTIC SEA<br />
3.1. Sampling: general aspects<br />
Miscellaneous processes, like seasonality, daily vertical migration,<br />
swarming, etc. produce typical zooplankton distribution patterns (Haury et al.,<br />
1978). In reality, we observe a result <strong>of</strong> <strong>the</strong> combination <strong>of</strong> those processes,<br />
and skilful observation strategies need to be applied in order to distinguish<br />
<strong>the</strong>m by choosing proper sampling duration (length) and measuring intervals<br />
(Sameoto et al., 2000). Appropriate measuring intervals are to be chosen<br />
considering <strong>the</strong> so called Nyquist sampling <strong>the</strong>orem (Nyquist, 1928).<br />
Following it, a signal must be measured in equal distances <strong>of</strong> more than two<br />
times within one period (or wave length) <strong>of</strong> <strong>the</strong> specific signal. O<strong>the</strong>rwise one<br />
produces aliased results as Figure 3.1 illustrates.<br />
140<br />
Measurements <strong>of</strong> a 24 h periodicity by different<br />
sampling intervals<br />
120<br />
Abitrary units<br />
100<br />
80<br />
60<br />
40<br />
24 h periodicity<br />
8 h sampling interval<br />
13 h sampling interval<br />
28 h sampling interval<br />
20<br />
0<br />
0 24 48 72 96 120 144<br />
Time [h]<br />
Figure 3.1.1. Examples <strong>of</strong> sampling intervals producing accurate (by<br />
8 h) and biased (by 13 and 28 h) results <strong>of</strong> a 24 h periodicity.<br />
Nearly accurate results will be obtained if a certain process will be<br />
measured at least three times within its periodicity or a wave length. For<br />
example, <strong>the</strong> twenty four hour periodicity <strong>of</strong> dial vertical migration demands<br />
an equidistant measuring interval <strong>of</strong> at least eight hours (Fig. 3.1.1). Longer<br />
distances, like thirteen or twenty-eight hours, produce apparent results <strong>of</strong> a<br />
thirty-six hour or even a seven day periodicity, for example. Continuous<br />
measurements would produce <strong>the</strong> most accurate outcome in both phase and<br />
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