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An Economic Assessment of Banana Genetic Improvement and ...

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IMPACT OF TECHNICAL INNOVATIONS IN AFRICAN AGRICULTURE 7<br />

in popular crop varieties, genes must be inserted<br />

into well-adapted host cultivars that<br />

are either conventionally bred or farmer selected.<br />

The second criterion acknowledges<br />

that genetic engineering <strong>of</strong> important export<br />

crops makes them vulnerable to trade disputes,<br />

regulations, <strong>and</strong> political lobbies outside<br />

their borders. For traded commodities,<br />

full market liberalization implies that shifts<br />

in agricultural productivity may not translate<br />

into lower food prices. For nontradable<br />

food crops, downward pressure is exerted<br />

on prices by seed technological change because<br />

they are determined within borders<br />

rather than on the international market.<br />

This observation relates to the third criterion,<br />

because the vast majority <strong>of</strong> smallholder<br />

farmers in Sub-Saharan Africa consume<br />

part <strong>of</strong> their food production <strong>and</strong><br />

many are net consumers. Both urban <strong>and</strong><br />

rural consumers in these countries will<br />

benefit many times more from the price<br />

decreases that accompany technological<br />

change than will those <strong>of</strong> richer countries,<br />

because food occupies a large proportion <strong>of</strong><br />

the budget <strong>of</strong> low-income people, <strong>and</strong> they<br />

are more responsive to price changes in<br />

terms <strong>of</strong> quantities dem<strong>and</strong>ed (Pinstrup-<br />

<strong>An</strong>dersen <strong>and</strong> Cohen 2001).<br />

Reduction <strong>of</strong> crop losses <strong>and</strong> yield increases<br />

can still lead to a significant increase<br />

in the income <strong>of</strong> poor households if<br />

markets function well. Focusing on the productivity<br />

<strong>of</strong> food crops, when bolstered by<br />

stronger markets, will also have national<br />

<strong>and</strong> regional income effects. Recent analyses<br />

confirm the potential role <strong>of</strong> food staples<br />

in domestic <strong>and</strong> intraregional markets as a<br />

source <strong>of</strong> growth in dem<strong>and</strong> for African<br />

agricultural products <strong>and</strong> income, in comparison<br />

with traditional <strong>and</strong> nontraditional<br />

export crops (Diao <strong>and</strong> Hazell 2004).<br />

In East Africa, the case <strong>of</strong> bananas genetically<br />

transformed for resistance to pests<br />

<strong>and</strong> disease meet these criteria. East Africa<br />

is the largest banana-producing <strong>and</strong> bananaconsuming<br />

region in Africa. This research<br />

focuses on the principal banana-growing<br />

areas around Lake Victoria in Ug<strong>and</strong>a <strong>and</strong><br />

Tanzania, which form a contiguous area<br />

stretching from the northeast portion <strong>of</strong> the<br />

lake, through the historical locus <strong>of</strong> Ug<strong>and</strong>an<br />

banana production in the Central Region<br />

(including the new locus with more<br />

market-oriented production in the southwest<br />

highl<strong>and</strong>s <strong>of</strong> Ug<strong>and</strong>a), to the Kagera Region<br />

in northwest Tanzania. The East African<br />

highl<strong>and</strong>s banana (EAHB) is the dominant<br />

type grown throughout this expanse, <strong>and</strong> a<br />

set <strong>of</strong> common biophysical factors constrain<br />

productivity. As the chapters <strong>of</strong> this report<br />

show, bananas are produced largely for food<br />

consumption, with surpluses sold in town<br />

<strong>and</strong> city markets. Growing urban markets<br />

contribute to the importance <strong>of</strong> the crop as<br />

a source <strong>of</strong> cash. Small quantities enter regional<br />

trade, with negligible exports. The<br />

population <strong>of</strong> the area is related linguistically<br />

<strong>and</strong> culturally, though separated by<br />

borders <strong>and</strong> administrative divisions, as well<br />

as by important political <strong>and</strong> economic<br />

differences.<br />

Though the origin <strong>and</strong> center <strong>of</strong> diversity<br />

for bananas is believed to be Southeast<br />

Asia (Simmonds 1959), the East African<br />

highl<strong>and</strong>s is recognized as a second center<br />

<strong>of</strong> diversity for an endemic genomic group<br />

(AAA-EA), which comprises two groups<br />

defined by their use in cooking <strong>and</strong> brewing<br />

local beer. The endemic group is also classified<br />

in terms <strong>of</strong> clone sets. All EAHBs are<br />

triploids. With three genomes, as opposed<br />

to two or four, few cultivars are fertile.<br />

Though pr<strong>of</strong>essional breeding <strong>of</strong> bananas<br />

began during the 1920s, it was not until recently<br />

that a major breakthrough was<br />

achieved through the development <strong>of</strong> a hybridization<br />

technique by the FHIA. Fertile<br />

male diploids are used to pollinate triploid<br />

cultivars to produce tetraploid hybrids.<br />

These are crossed again with improved diploids<br />

to give secondary triploids, from which<br />

selections for release to farmers are obtained.<br />

However, the endemic cultivars most<br />

preferred by farmers are infertile <strong>and</strong> cannot<br />

be improved by conventional means.<br />

This difficulty <strong>of</strong> improving bananas by<br />

means <strong>of</strong> conventional plant breeding tech-

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