Psyiiids (Psylloidea, Homoptera) of the Canary Islands and Madeita ...

81 1976
I
Psyiiids (Psylloidea, Homoptera) of the
Canary Islands and Madeita
M. M. Logioova
Zool. INUNU of me Acadcmy of Scicncn. Lningnd. rhc USSR.
Societas Scientimm Fennica
Heisinki - Helsingfon

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Copyright e1976 by
Socic~r ScienriuUm Fcnnica
ISBN 911-613-054-0
ISSN 0069-6179
AbStnet
LOGINOVA, M.M. (Zool. Institue. Audany of Sckncc. Lcningrd. USSR): PsyUidr CPsyUoida,
Homopiem) of the Cpiury isiandr and Mdeha. Commentur. BioL 81.37 pp. (1976).
This paper u the fint to contah a suumy and dcsuiption of the psyUid fauna from the nonhern
iriandr of MacPronaia. AU the 29 rpecies b td below ate na>rded from this tenitory for the
fmt time.
Authw'r uddmss: DI. M.M. Loginova. 2nd lnrtitute of the Acadany of Sciencu, La@%!
8-16), the USSR.
. .
.. -
i
In this study 1 made use of the collections of Prof. Hakan Lindbcrg, who visited the
Canary Islands and Maderia several times (1926-1963). This paper, written in
respectifui memory of Prof. Lindberg and dedicated to him, completes my dysis of
hk mcxt interesting collections of the Psylloidea fauna from north-western Africa, a
temtory which up till now has been p d y investigated as rqards this group of
insects.
The species dealt with hen inciude psyliids coilected by Dr. Remane from Temrife
in 1954. These collections were sent to me for investigation by Prof. W. Wagner
(Hamburg), to whom 1 edpress my sincere gratitudc.
Description are given her of two genera and 20 species up tü1 now unknown to the
scientific wodd. The holotypes and paratypes are in the keeping of the Zoological
Mwum in Helsinki (Finland) and of the Zoological Institute of the Academy of
Scienees (Leningrad, the USSR).
The mcasurements are given in millimetres, and ihe abbreviations in the
mcasurement list are the same as in the author's paper on the Psyllids of Morocco
(LKINOVA, 1972). The geographical names are talten from the original labeh. '€le
collector's name is given when the material was not collected by Rof. Lindberg
himself. The Qures are original. The same part of the body and details of its otnictun
in members of the tribe Aryfahhi are shown at the same magnifiutionr, so that both
the distinguishing morphological characters and the tme &e of the species may be
easily grasped and compared.
Aph.IPridac
EuphyUura Foerster, 1848
Reprrrntatives of thir genur from thc Oid World show very nmw trophic speculizrtion, bei
resuicted to OIca L and Phü'yreu L. (fam. Olucue), and have a unifom st~cturc The speckr ue
dUthyished by the fonn, venition and psttrm of thc fonwíngs and the mile gcnitaiia.
Two rpeciu have becn discoved on the Cuury islands, E.divin0 (Costa), huc oecurring at the
wutemmost point of its range, anda ncw rpecier - &.conarie~~& Lo&, describcd in the mision of
the tribe Auphyllurini (Loginova, 1973).
HUSINGlN WKUSiüüJAPAiNO OY
HUSlNKl 1976
Euphyllum divina (Costa) - Gran canaria, Tafiua. 24 d 9, ieg. R. Stort
On spccks of O&u. in Europc on O.cumpueu (Loghovi, 1972:4).

Euphylium CIVILV¡UI& Loginova - Tcnerife: Vaik S. 2 111 1950, 3 dd includin a
hoiotype. 5 99; A-. Bailadero 700 m. 2 I I I ~ W e s 28-30 , V 1947. 2 69
(on Pmmn iusiraniru~~'Gran Canaria. S v.lar Palmas, 5 111 ¡SSO. I d.
Fom and venation of forcwings like thore in E.o~hina, but even more ¡*e thore in€.oct&iopua
Silwstri. 1934; ptcmrtigma without transversal vcins, ükc that of Gacrhiopicu. apcx of dorsal platc
in femalc turncd downwuds, genital scgment in male quadrangulu (lateral v~ew). Thus.
8.canarunsU rcprcxntr an intermediate specier bctwetn two morphological g~oups of the
palrcuctic specicr of thc gcnus - the grwp Phülyieae, which ir chuicterized by the lack of
tnnnersal vcinr on the pterostigma. and the ~IOUP Olivina, having a longer pterostigna. croswd
with transversal vcinr
Agonoscena Enderlein, 19 14
Agonoscena tugionii (Licht.) - Gran Canaria, Tafua. 8 dgcieg. R. Storl). On Pisrucia lenfiscus
(Loginova. 19721).
Srrophingia Enderlein, 19i4
nie typc specks of thir genus. SericoC (Curtir, 1835). is widerpnad in Europc, extending
nomvards to Central Finland and thc Leningrad redon in the USSR. and living on &#una
vulgatis. Tht rccond spccicr of the genus. S.cinemac Hodkinson. 1971, from England. tivu on
Enea cinerea
Two new spccus are dcscribed below. They werc discovcrcd in the mountains of Mideira and
Tencrife on arboreous hcathcrs. mostly €rica arbma, which suggests that thc Snus ir confmd to
huthcn of the subfamily Elicoidco.
Sfmphin#ia arborca Loginwa, p.nov.; f i 1-11.
Dcscribed frorn a series of 31 spccimenr ftom Mdeira and Tcnerife.
nie cmaging inítctr arc green, the predominant coiour of the pechens from Tenenfe.
Howevsr. Iatei on thc body bccomer yellow. the change beginning with the head and ley and thcn
anbracing the thonx and abdomen.
The mijority of the specimens from Madcin ue ycllowish. having no patlun on rhe dorsal pirt
of the thorax; thc tarsi, antennal regmentr 8-10, the apicei of the genitrlir in the femak and thc
puamuer in thc malc are bmwnish to bmwn Eyes blackish or fulvous-biack. Forrtvingj more or
Icír intcnr ycllow. xidom beuing a faintiy deficd, pak brownish coral-like pattern. mostly in
ccll rs, veins light-colourcd.
Body roburt owing o wide. convu, dom*like thorax and short appieued had. The anterior
put of the scutum of thc meronoNm reprcrnts thc apu of the uch fonned by the hud and
thorax (in lateral view). Hud nurower than thorax and preued to it. pmplcuritu gcncdy
invisible. Vertex even, straight poiteriorly. slightly convcx anteriorly. changing gndually into
genac. Fweae shdlow. Daucd occlli on lateral mugins of vertcx, at its hind angics, unpakcd ocellus
on a very small crimson rhombic frontal sckrite in facial plane of hcad and hardly viribk from
above. G e ~ not e swollen. Pronohtm rectangular. its siightiy narrowing sides curled on to pleurd
part of thorax. almost hdf as long as pnescutum of mcronotum. Fommgs in both male and
female cggshaped, Costa anylarly cuned in cell c; with a tapcring roundcd apu; no costal break;
ptcrostigma disiinct, tcminating behind middle of ccll r; cells mi and cu mirlt, more 01 lesr qurl;
vcins not strong. Manbrane EO~C.CCOUL with few tranrersai wrinkies, aimost opaque.
Hind wings romewhai shorter than fonwings, mcmbranous, tnnsiuccnt. widest at the roundcd
apical mqin: M and Cu - onc-branched. stem MCu short.
Lcgi short with shon strongly dLted femon: mxae oí hind legs with a long mcnunt, tibiae
basally smmooth. with distinctly düated apiccr having 7 equaliy spaccd saltatorial spinez Fust
agmcnt of tusus with two such spincs
Fcmak. Geniialia cuncatc. fairly murive. basaily thick (fig. 4); dorsai platc slghtly cancave
rbove. Ventral plate basaliy aimost twtc as high as dorsal plate. Extcriorvaives wide.
Figx 1-11. Snoph¡@ arbwm Lo&inov& p.n.: 1 - hud. from abovc and below, 2 -
antenna, 3 - forcwing of male, 4 - genmltrlir of fanalc. in laterai v h . 5 - gcnitrlir of mak, in
latd view. 6-7 - paramaes, from inride uid from abovc. 8 - 2nd segment of p ea 9 - htid
leg. 10 - apcx of tibia and tusur of hind kg, 11 - es.
Malr Ami tube with a faintly dkicd CIIV~ posterior mar@; about 1.5 her as h@ u
parunaes and genital xpent Panmercr shple, widut at bue, ttpering at apu. Penh kiit
downwdr at itr wide opa, ductus short. riluated above pomt whm apa bendr dowmvudr
Egg w. narmw, with a shirpcned anterior pole and a thick attaching rtmi at ¡U porwior
pole crí 11).
Meuurcmcntr of femalc: kngth 1.92-2.50; kngth of fornvings 1.62-2.02. width 0.62-0.80;
width of hud 0.55-0.65. width of Vertu 0.39-0.42. leiyth 0.22-0.25; l~iyth Of UIkWUC
0.30-0.37; avcr8gc kngth of dorsal phte 0.50. Mcuunmenta of mak lcngth 1.67-1.87; length of
foiewm 1.30-1.50, width 0.55-0.62; width of had 0.50-0.55, width of vertu 0.32-0.37.
length 0.20-0.22; lcngth of antennae 0.30; he¡@ of pmeret O. 12
Madein: SMaddcnn. 4-5 VI1 1957. 1 9 ; Quehadas. 24-26 VI 1957. 2 99. 14-16 v

6 M. M. Locorovr. PIyllids of thc Cvwy LIpD

LiYonia echidna Loginova, V.nov.: f i 17-30.
ikrribcd from 8 males and 3 femalea from CranCanuia.
Body light-yellow. fovuc. dents on thc prOnONm and apex of paramena browh Eyes yellow
or brownish diver; antcnnae dirty yellow. rgmcnu 8-10 brown Forcwing mcmbnnous, but
fauly dcnr. aimost opaque. faintly yellowish. Hind wings membranous, colourley exccpt for the
yellow clawr
Vertcx slightly longer than half its width; paued ocelli on lateral sider of vcrtex. and a seta on
cach mar ande. Unpaired ocellus invisible from abovc Svellings behind the eyes widc espccLlly
neu verla. Csnac faintly dhted. clypeur lying in a shdow dcpression behveen than. distinctiy
visible in anterior view with 2 long rtae on its anterior muga
Proború compressed bctween coxac of fon kgs, itr long terminal put direeted fotwudr
Antennac aImost as long as width of head, thc rgmcnu gmdudy thickening from 3 to 8. but
rgmcnti 5 and 7 thinner and shorter than thcir neighbourr Tibiae of hind lega with 5 qudly
spaccd vltalorirl rpincs, brownhh 01 light-colourcd; mcracanu having the shape of a md obtuse
Nbercule.
Fonwingr Ion6 rounded. with apex in ceU rs veins rmngly raised. the nodal lme abrnt. M
bnnching off at dirtinctly shorter distance from base of wing than R+ Ptcrostigma wide, long;
thcn ir a costal bnak. cells m , and cul luga.
Female. Gcnitalia longer than other segmcnu of abdomen, dorsal plate concave above, ventral
plate slightly ninower than dorsal plate and half as long (lateral view), iu apex rectüineu, so that
the broad cxtenirl vrivcs of the ovipositor are free.
Mde. Anal tube convex anteriody. dightly dbted porteriorly. Pumercs somcwhat lowcr,
tapering and rounded at apex. termimting in a narrow swelling turned inwudr; the geatcr put oí

10 M. M. LOCINOVA. Pqiüds of

COMMENTATlONE3 BKuw;icrre 81 13
bcnt Corta in CCU c; veins faintly raid above mcmbm, rtcm RRr dihtcd. Rs very bng. gently
S-rhaped in opid hU. pteroitigma terminating bchind middk of -11 r. R dirtinct thmughout itr
kngth. Thc brinching off from the root of thc wiy ir disphccd towud, the base id oaun in thc
main third of itr Icngth. stan RRs U approximately quai in icngth to MCu and the root of the
wing In all the celli the mcmbrane ir cwered with minutc thkk superticid q k r up to the vcinr
Hmd wings mcmbmus. the cianu alom ir dense; branches of M long here u well u on Ioming.
Position of rhinuia on antcnna imposible to dhrn without miuorcope.
The genitah are shown in Figz 36-39. lky are very thick in thc femdc, markedly iniaccd in
the dorsal phte bchind thc Uolitcd and rcgion
Anal Nbc of male convcx antcriorly widened portcriody in the rhrpe of a biade. with a
triangular protuberance at the ex; it u twh aa high u the genital scpent "he puamenr
rcetanguiuly Ismcllatc, thh, with trianpiar projectionr in the lpral third of the poatssior mugin
(dorsal visw).
Muumncnt of femdc: kngth 3.65; length of forewinga 180. width 1.0; width of hud 0.72.
width of vertcx poatcdody 0.40, anteriorly 0.52, length dong epicnnirl auture 0.35. with lobe;
0.52; kngth of ante- 1.45; length of genitoln 0.87; height at base 0.57. Mururrmenta of male:
kngth 3.0; kngth of forewings 2.32. width 0.85; width of hed 0.66. widths of vertex 0.35 Ud
0.45 rerpcctively: kngthr 0.32 and 0.50; le@ of antcnnae 1.40, height of uul bubc 0.37, of
paramuu 0.30. of genital sepnent 0.20, length of genital rcgnent 0.30.
Gran Canaria: Buiduna, 5 IV 1%3.1 d a holotype; Tafua, 1 9 (kg. R. Stod).
"he spcekr and genur derribcd above undoubtcdly bcbng to the family Apliobrtdoc nK
unque rt"cture and lpck of any biolo&d infmation make it diffmlt to idcntify itaplcr: in thc
family. Howeve~, aich spe& luaimr as the strongiy developcd fon lober pad tlatmrr of the
vcrtcx. dense rhombic fonwinga with long M baiichcr and genital stiucture rugcrt a connection
bchvccn Megudkmnia pn.n. and Ihe genur Euphy/h Frst. It ir probabk that thc new gen= ir
endcmic to the CÍnary IrLnds.
uaiaily bcing pdc. Dod pattcrn Wre that in A.gen&fue Latr.. dark brown; thonx klow, feanon
of Iq, and abdominal rletiter hnhg ths me
qota, whwi Liaoue with qo. Fbt Uuw .Iugmcnta
yellow browr the colair dulcning progedvely bi the fdowin# aepwntr Eyes
crimrobfukous, occlli redduh. Facial and cervical put of hud, ki m e cuca incLding the
clypeus blck-brown
PSyW8C
Diapbrina LOw, 1879.
Dbphwh ContuUrO LO&- Lamote: Oir~h. 19 111 1963. 1 d; El G ~ O ,
Fuerteventure, Martural. 23 111 1%3.1 d. 1 9. Islas Cpnvis. Lobq 28 111 1963.1 d.
Hort plant unknown ücrribed from Morocm(Loginaa. 1972; 15-16).
16 111 1963,25 d9.
Atytaina Frst. 1848.
Representativer of thu genus are disiributcd di over the world in regions with I subtropical
ciimrte mortly occunuig on lNbs of the Leguminosac. More than 30 pecics havc becn reported
(HeilopHaf~iron, 1960), but in my opinion thc genur is hctcrogencous and in urgent nced of
mirion
The main chuactcrr diagnostic of the pnus Arytaina:
i/ exterior wltatorial vine on furt ugment of tusus of hind lcgs.
21 vey wide hcd (wider than thorax). with sphcriul eycs.
3/ gmduil tranaition fmm vcrtex to gcnd cones, which are not contipous u thcii baacs ncu
the unpaircd occllus,
4/ Iack of pterortigma on the forewinga (in somc cases it is Ndimentuy) and of costal break,
51 rtrong thorn on base of hind tibiac.
%o ncw rpecics and onc subspcaes fmm thc Cuiory Ishndr are dercribed bebw.
Arytaina mtbwqa Loginwa. v.nov.; fuk 40-50.
ücrribcd from 136 spccimcnr from Tencrife.
Body yellow, coloui varying in intensity betwcen spccimens. somc onnge or rcddirh, the hud
Figi 40-50. Aryiaina nubffqp7 Loginwa, va.: 40 - hcad, from above and bclow, 41 antenna. 42 - clypeus, 43 - forewing of mdc, 44 - geniWu of female, in iatcnl vbw, 45 -
genitalia of mal+ in lateral VKW. 46-47 - puamercs, from inride d in antctio~.intcrior vkw. 48
- apcx of penis. 49 - hind leg. 50 - egg.

1s
Forewing colwrleu. scldom faintly whitish, having yellow patcher in centre of uch ceU
exccpt ccil c íFiS 43); thcrc prtchu often diminishcd to smrli "tonycr", from which rk a series
01 minute biown marginal rpincr Veinr dark brown
The cdontion of thc body, $Le and rmcturc ue very Iike thoít ofA.gcnistue, but the hud is
much widei than thc thoru and slopa downwuds, thc facial piane u widcr and thc Iiom U almost
mund. The trirnplu genal coner are apWy httcncd Ud tum downwuds. whereu in.
A.gcniutue thcy are parakl to the wrtcx. ll~~
wrtex has dccpcr foverc. its posterior angles are
faintly dhted and incline downwards, projccthg in thc form of rounded loks, uch of which has
an occllur in ita centre. The unpWed occüuí U invisible from above, since the fore loks of the
vertcx are contigous; the anterior mara of the vcrtcx ir gcntly archcd, but not trianpiar like that
of A.genistuc.
Forewings narrow, with anterior and posterior margins more 01 les paralle4 widcly rounded
apkaily. 3 times u Ion8 as their width; pterortigmr Iacking. but the region whcre R fusines with
the costal mara dightly dhted. SupcrfKkl spher in cell NZ around the anal íuture, marginal
q~inu foming nwow üner in cch m 1. m2 and cu 1.
Ccnicrlir of femile as in A.gcnistue
Ceniiilii of mok differcnt in ccttain detlür: anal tube eithcr higher 01 as high as genital
rgmcnt at ita bar; its protruding apcx having the fom of a wide Mg. The genid tegmcnt ir kger
than in A.genirtue. and extended longitudinrliy. The parameres have a different outlinc, with a
l~tudinrl weliiig interiorly, ao thit they are contipous at two points on the apa, one below
the other (iir 45-47). The penu rernnbler that ofA.genistue.
Muumnentr of femile: length 3.5-4.0; length of foming 2.9-3.25. width 1.0-1.2; width
of head 1.05-1.12. width of vutcx 0.6-0.67, length 0.25-0.3, len@h of gcd wncs 0.15-0.2;
kngth of antcnnac 1.95-2.22; kngth of geniwk 0.75-0.85. biurunmcntr of nuk: length
3.3-3.62; length Of fünwiryr 2.72-3.0, ~idth 0.95-1.12; width Of hud 0.97-1.1, width of
vutcx 0.55-0.62, length of pn.1 cones 0.15-0.2; length of antennac 1.97-2.22; average heyht of
anal tubc 0.37, genítrl tevent 0.35-0.36. hewt of panmms 0.20.
Tende: Pico di Teyde. 36 d? (kg. R. Sto&, 53 d?, Muding 8 holotype d (kg. R. Fny);
Teyde, Canada 20 V 1947.9 d ? (oa Spurtffyliarr nubbnur and Cyhs); Las Cañadas. 14-15
11 1940, 18 d? (on Spmocyfinrr nubbenur); Las CaWas 2300m. 16-17 IV 1950, 8 dv ;
Canadas 2300-2500 m. 28-30 III 19S4, 5 dd 0%. Dr. R Remane); Lu Cliuny 3 IV 1959,7
d9.
Both the charactus similar to tharc of Agenistur and tharc diffcrentlting Anubhugu are
cmphisizcd in thc deiuiption. The new pccicr wu collected on SpartffytinrZ nubigcnus and
Cyfiorrrp., which are probrbly its host plan&
Alyah drviu Loginovp. sp.nov.; fiis 51-63.
Dcrrikd from 87 pecimcns from Tcnerile and Fuerteventura.
Young spccimcnr geen 01 iight yellow. pattern on herd and thonx yellow to onnp. Md
cpicrankl íuhlre, and facid and ccniui puta of hud brown. Cdour darkeniq with igc; pattern
on upper put of body becoming brown. and Iowu part of thonx. tibue of kgr and abdominal
sckriter becoming covercd with bmwn spots, so that imaUy only n mw stnp bctwecn pattern
on hud and thorax, gend coner and manbrane betwccn abdominal sckriter remrin unpotted.
Figs 51-63. Alytuinu d da Loginova. rp.n.: 51 - hcod, from abovc md below. 52 - antenna,
53 - clypeur and proboscis, in Iatud vicw. 54 - proboscis. from below. 55 - fmwing of femrle,
56 - genitalia of female, in lateral view. 57 - geniWia of male. in laten1 vicw. 58-59 -
puuncres, from inside ind behind. 60-61 - paruncres, from behind and above, 62 - apcx of
penis, 63 - hind leg.
Figs 64-66. Arytuhu devio insuhrir Loginon. subsp.n.: 64 - genitrlir of mak, in lateral VKw,
65 - gcnitalir. in posterior vuw, 66 - paruneres, írom ibove.

16 M. M. Locarovrr. Psyllids ofrhc Cauq kLiidt d Midelln
Forewings coiourieu or palc yeiiow, fairiy oftcn with yeliow patches in centre of ccils, Umilu
to thor in A.nubh@ sp.n.. but snler and paler. Vcins brownuh with dUtinct dark brown spotr
at apicer, and on marginal vcin under a series of mmute marghd spkr Antcnnac. cyes and ocelli
coioured iikc thora of A.nubhaga
A.&& combina the morphologiul charactsls of A.grn&&e and A.nubivugu The hcad U Wre
that of A.genisiue, the vcrtcx triangular antenorly, the fiQd phe betwocn the vcrtex and short,
almost contiguous. genal canes U narrow; thc unpaind oeellur U visible from abovc However, thc
heid ir not wider than the thorax, which resanbks that of A.nublvqpr. but U mon convcx above;
the rutum of the meronotum ir iongcr in rclation to the rizc of the pronotum and pracrrutum.
Forcwings diitincily widened towardr the apkal third (of the whoie Icngth) with avwy wideiy
rwndcd apial mugin, which u not typicai of the rcprcrantativer of the gcnus; Ndimenlaxy
ptcrortigma, fonncd by the diiaüon of C and R, which is not alwayr strongly mukcd, cell cu1
high; no minuto aiperíiiiai sphes.
Gcnitaiu of femalc as in A.nubhagu, but narrower at apex.
Mak. A d tube Wre that in prccediag specks, genital rgment shorter, llmost sphericll. iowcr
than anal tube. ApW blnde of pcnis, tumed downwardr, shorter and widcr than inl.gen&tue and
A.nubivuga. Thc parameres compliuted, hteraily ruembiii those of A.nubivau and porteriy
thor of Ay//un&rita (Zctt.), Uncc thc posterior margins of the pacamere lunellre uedüated and
fonn widc bbes, denrely cowrcd with small rtac (fig. 58-59).
Murumnentr of female: kngIh 3.1-3.5; lcngth of forewingr 2.6-295, width 1.12-1.17;
width of hcrd 0.77-0.85. width of vertex 0.45-0.47, kngth 0.2-0.22, kngth of pnal conw
0.07-0.1; icngth of antcnnae 1.42-1.75. Muinments of male: kqth 2.8-3.1; kngth of
foccwings 2.35-2.55. width 1.0-1.05; width of hud 0.75-0.76, width of vertex 0.45, kngth 0.2,
Icngth of genal cones 0.07; lcngth of antenm 1.55-1.52.
Tenerife: Valle de Santiago. 12-13 V 1947.13 do; Aqua Mansa, 15 V 1947.2 99,l d and 8
II 1949. 1 d; Teydc. Canadas. 20 V 1947.41 d9, a hoiotype d among them; Bermejo 1400 m,
21 V 1947. 1 ¿ and II II 1949.22 d9; Las Cyíadas 14-15 II 1949.2dd; VMor 1500 m, 27
11 1950, 3 99, 1 ¿ (on Cytisuspmiiferus. AdenowpusvimsusJ. Fuerteventure, LaOüvo. 16 111
1949,19.
A.devm v.nov. differs from ail thc known representatives of thc gcnus in the IIUTOW hcpd, the
forewings, widcning towards the apex and hiving a NdhenW ptcrostigma, and the dody more
convex thorax. Ther features are ahared with the rcprcrantatives of the pnur ArytlinillP and
tcstify to a genetic rciation ktwecn thue two genera. A.dcvfo sp.n. re~scmbiesA.nubivugu and has
ken collected from the m e stations in a numkr of placer
AryiaiM devia insularir Loginova. subsp.nov.: ti@. 64-66.
ihe rue. sttuctum and coloration of the body are iike thosc of A.ddu sp.n. ihe structure
of the panmeres u dutinciive; the posterior marginr of the paramcre plates are only dighliy
widcned at thc bw, whüe in A.devm thcy have iugc rectangular almost cintiguous bhdes (cf. figr
60 and 65).
The rpecimens on whkh thU aibrpecies was separatcd were found on the W s of Paima and
Gran CuurL, where the nominal species was not discovered.
La Palma, El Paso, 6 VIII. 2 dd (kg. R. Firy). 26 V 1947, 1 d. Gran Canaria: Valle de Tejcda,
7 VI 1947.1 d; Santa Brigida, 21 11 i949,l 9.
Avtainil& Ramirex-Comez, 1972.
For the nomencbtbc explanation regarding the denomhation. compodtion and diagnosis of
the genus, LCC my paper: Loginwa 1972: 17-18. Thc materia¡ collccted by Prof. Lindberg in thc
Canary Idandr has inuoduccd 10 ncw mcmbers, so that the genus now comprks 18 specic$.
lk rpcckr from tbe Cuiuies iad the two rcpnantativer fmm bíomcco ampose a domi
group of pnstially nhtd rpccia, cmrized by a seria of common morpho~
features:
short, blut genal cona mt widely ap8rt; wide. pcatago~i, bus fmnt, unifonn atNctum of nuk
gcnitdia Howcvei. them 12 lpecks us clurly divided into two morphdq¡d wpi. mosW by i
diffennce in the structPn of the funde pnitrlb
m fit group amtuna four exclurivcly insular rpccWr: A.pboreidu. A.n&whm&
LdivLlcnr and A.rnudim spp.a The fanale has iong, murive the body b iUs and the
proborb considerably bqer üua usual
nic sccond pup mntillu six rpecier fra the CanaricrA.&luio. A.quilonc, A.p¡k&&,
Aprognatu, A.iruirbu and A.umhtu spp.n.. and two morown rpccks, A.hakun1 b& uid
A.colF.r. Lw The chancten &tinyiJiLig thus spcckr M: rel.tivdy short pdWL in the fsm,
le. with more 01 iur rtmigly deveioped bldbiikc projcciionr it the interior margbr of the venW
platc. theu aiillcr size iad a proboscia of more uaud lengih.
ihe remlinmg six representativa d the genus have no speciai peailiuitier terti@& so súoogly
to thcir morphdo&ai unifomiity; their origin aiso differs fra that of the abova

antcriorly mered with rtae. The proboscU is often vcry long. projecting fu beyond the anterior
mar& of thc head (latual view), however in the mjority of the rpecks it U of more unid length.
The elongaiion of the proboscu should appuently be considered an adaptive phenomenon,
depending on thc depth of the vcucls from whkh the insccts are sucking and on the ande at which
the hud of the insect U inclincd when it U stabbing the tissue of the plant.
Representativcs of othcr genera of the tribe Arytohini show a similar kngthening of the
proboreir Amblyrhina cognoto LOw has a proboris as long as that of Aryfohilb proborcideo sp.n..
whüc in a rclatcd speckr Ambiyrhino mculnfo Low, the probosck U ffen mewhat shorter than
usual. Thb suwstr thit the lenglh of the probosck has not thc tuonomic vrhic attribtcd to it by
HerlopHumoh
Thonx mioothiy convcx. pronotum approximately twke as wide as length at sides, postcriody
aipcrimpord over the pracmtum of thc mcronotum. whkh is relatively longer in thc species of
thU gaip than in the other representatives of the genus, praexuNm * twke as iong as pronotum.
whkh U bent forvuds and downwards
Forewhp manbranais, widest in thc apkal third of the whole Iength, wiih a widcly rounded
apicpl mar&. pteroítigma WCU dcvdoped. denser than wing membrane. bordered intcriorly by
vividly muked R.
in the furt four rpecics, thc females have long massive genital¡; in the remiining six new
spccecu>. thcy havc short. more or lus conaükc genitalia.
Arytainillo proborcideo Loginova. sp.nov.: fyrr 67-77.
Dcrribcd from 59 spccimcns from the utpnds Tenerife. Comen, Gran Canaria and
Fuerteventura.
In thc oldest spccimens, besides the general yebwing, thc amcd Ud facial puts of the head
and the anterior put of the pronotum have become bmwn-blck; thc abdominal steinites ue oftcn
bmwnirh. Anlcnnac duty ycibw, scgmcnts 4-6 bmwn at apcx, rcemsnu 6-10 whoUY bmwn.
Forewinga hy.l& with nmow yeUm prtches in SUS of apical half of wing and around anal
aiturc; the pitcha may diminirh to mmute lime& from which u*e a scr¡cs oímarginal sphiu; veinr
duk brown, bordered with shart lightsolaircd dashcs Superficial spines dark, occupymg only
central part of ceilr. kking in ceU c.
Femalc. Ccnitaiia mrírne, long, strktly ainute.
Mik. Anal tube ahost undüatrd postcriody. slightly hucr than paramms, and genital
-ent. whkh h widely roundcd Wow. Paramenr with a dktuict bi.de along anterior margin.
whkh U coverrd with long dense rctu. Nmw triangular apices of paramerca touching only at
theu posterior andes (view from above. fig. 75).
Meuurcments of lemale: kngth 2.87-3.60; kngth of forcwings 2.37-3.07, width 1.0-1.25;
width of hed 0.75-0.9. width of vcrtex 0.42457, kngth 0.15-0.18, length of genal ames
0.04-0.06; kngth of antennac 1.5-1.7; kngth of dorsal plte 0.92-1.37. Mcasunmcntr of mik:
kngth 2.8-3.15; bngth of fowings 2.5-2.62. width 0.95-1.07; width of head 0.77-0.85,
width of wrtex 0.45-052. length 0.15-0.17. kngth of ged cones 0.05; icngth of antennac
1.52-1.62; hejht of ul~l
tube 0.32-0.37, hejht Of PM~CES 0.30-0.37.
Tenuife: Pico di Teyde, 2 99 O-. R. Ficy); Teyde. Canadas, 20 V 1947. 20 d9 (on
Adenocorpur virconrs); Las Cañadas, 14-15 II 1949. 19 d?; Las Cuudas 2300 m. 16-17 iV
1950, 1 d. 2300-2500 m. 28-30 III 1954, 12 d9. inchiding a holotypc d Ocg. R. Remane);
Viflor 1500 m. 27 II 1950, 1 9. Comen. Hormiya, 1 d O-. R. Frey). Cnn Canaria, Moya, 1 ?
(kg. R. Stod). Fuerteventura, 10 111 1949. 1 d.
A series of 5 fanaics from Tcmrife (Agua Mana Porao, 2 V 1947; lns Canarir. 3 V 1947)
from the colkction of the Rijhurum van Nituurlgkc Historie in Leyden (Hohd), which wu
scnt to the author for identirication. is also designated as paratypiul
Arytohillo proborcideo sp.n. rescmbla A.n&oIinmla sp.n., dcscnbcd below. but diffcrs in its
forcwings whkh are more widcly roundcd at the apex, lack supcrficll spincs in ccll c and have
F&s 67-77. Arylohüla proboscideo Loginova, q.n: 67 - hud, from above and baow;68 -
ilyilum of antenna. 69 - clypeus and probos& 70 - probos&, from below, 71 - fomwing of
female. 72 - gcnitaüa of female, m lateral vinv. 73 - geniWY of malee, in htcralvicw; 74-75 -
parameres, from inside and above; 76 - apcx of pea 77 - hind kg.
strong duk veins with bght-colaired dasher; the form of the puuneres U also diffeicnt in thae
two rpecrr
Host plant unknown; it may be Adenocorprr vkcuw from a tcrier of insccts ms collected
Arytoinilia nipalineata Loginova, sp.nov.; TÍr 78-00.
Dcscribed fmm 46 spccimens from the idandr Comna, Gran CnauL and Tenerife.
Brown pigncntation not uually apperuiiy when the body becomes yellow. Fomwing
tnnducent 01 whitish. ycllow patchu in thc aUs dimmer and shorter than in Aproboocidcn.
Difiers from A.proboscidea in thc very smiU downwards t u d pnal conas and in having 2-3
pairs of long dark setae on thc facial planc of thc hud.

,
!r , E -
20
21
1
genital rcgncnt Paramuer. withait biadoüke projsctions anteriorly, nurowing gradual& towardr
apcx. whue they fonn wide triui

22
I
CQWIENTAllONES Bwxociw 81 23
A~loinUb modka LOgnovr, rpsov.; í&r 90-103.
Dcscribcd fmn 9 mda Ud femiltr from the irLndr PJma lad Hiena
In iu coloration, &e and stNciun gratly nambh A.dlrldrnr q.a. from which it chbfly
diffur in the rtruc~~rc of the gcniwh. erpefiüly thois of the mala
Femalc. Dorvl pkte nrnowed, erpefLUy ilmg intefior m8rgin. in the pha whcm it puwr
into the ipicil thiid of the length. Intcrku muginr of ventnl phtc widcned in rhipe of long and
mow zigh t-anglcd bladek rtretching on to Mes of d d phte.
M e Puunuu pntly tapcrhg from base towuda blrde in anterior mqh, bu1 strongly
mowing ia upper part, u) that they fom bnnchcs, contipour at uitulor mqhr of trbnphr
rpeg drawn de. quite diffuent from in A.dwi&ns (fw 101-102;vkw from above). PortuiOr
@r of the piromem undilated. thcir interior mUpns thur having the outhe of an obbng oval
in portenor vKW; in A.diuAdenr thc lowu port of the ovil u conridembly MXIOWC~ owag to the
diiation of the posterior marginr
Muainmentr of female: le@ 3.3-3.75; length of forewings 2.82-2.97. width 1.1-1.15;
wylth of hud 0.85-0.87. width of vertex 0.5-0.51. hgth 0.21-0.22. kqth of @ CO~W
0.07-0.1; icngth of antclu~~c 1.77-1.9; kngth of dorsal phtc 1.0-1.05. Maaircments of mrb:
iength 3.0-3.25; iength of fonwingr 255-27. width 1.05-1.07; width of hud 0.8-0.82. width
c
.b
Figx 89-97. Arytainilh dividenr Loginovq sp.n.: 89 - hcad, from above uid below. 90 -
forewing of mde. 91 - hind wing of male, 92 - genitalia of femaic. in lateral VKW. 93 - genitalia
of male, in lateral VKW; 94 - paruneres from inride. 95-96 - apex of penk 97 - parmeres,
from abovc.
Figx 98-103. Alylainüh modh Loginova, sp.n: 98 - herd. from above and bclow, 99 -
fomwing of male, iüü - genitaiia of malc. in hterdview. 101-102 - puunercs, from inridemd
above. 103 - apex of pcnir

of vertex 0.45-0.5. kngth 0.16-0.2. length of genal cones 0.07; length of antennae 1.62-1.77;
heigth of anal iube 0.4-0.42. heigth of parameres 0.5-0.55.
La Palma: Caldera, 25 V 1947. 1 d; supra El Paso 600 m, 4 IV 1950.1 d, 1 9; supn El Paso.
El Pindi 1300 m, 1 9. Hierro, Guuuou, 26 III 1950.3 dd. including a hoiotype, 2 99.
Found on Palma with A.dividens, but evidently more common than it on the island. judgimg by
the collectionr Hori plant uncertíin.
Aryiuinüb diluia Loginwa, sp.nov.; Ti& 104-112.
Desdbed from IS males and fmaler from Gran Canaria Ud Tcncrüe.
UliuUy uniformly strawyellow. the dorsal pnern Lclting, forcwings colourlcu or whitish with
brown veinr; faint yellow patches sometimes found in centre of cells, crpecLlly beride a Iuier of
marginal spinu Firrt 4 antennal segmentr dirty ycllow; 3-4 apicdly, 5-10 wholly duk-brown.
FomvLyr widert ki.ond fust thud of their lcngth, minute supcrfkhl rpiner miss@ in ccll e;
in the res1 of the ceb iky approuh the veins. but lave wide rpuwlcrr bordcrr
Femilc Donrl plia wids; slightiy clevated in uul puf itr top süghtly bcnt upwdr Ventral
piate basaüy twyC as he u doraal, considenbly Bata, with bldbüke dilations ilq intoikr
marginr
Milc Anal tube WIOIY, porterioriy faintly düated up lo hilf itr heighf lower than puunt%~~.
Panmeres simply lamdiate, widenky at buc and apuc. rpially fomb a hook-iikc foiwud
cuned branch (vicw froi above. fi 110).
Merruremcntr of f d : kngth 242-275; km of forsaring 2.02-2.1. width 0.9-0.97;
width of heid 0.62-0.76. width of V ~ C X 0.37-0.47, imfi 0.15-0.17, lS@ Of @?IIAl COMS
0.05-al; kngth of antcnna 1.25-1.45; kqth of dorsal plite 0.65-0.75. M ~ m c n tofmik:
a
icngth 225-267; lcngtb of forwings 1.87-2.2; width 0.75-0.9; width ol head 0.61-0.69. width
of vertex 0.37-0.42. Ln(th 0.12-0.16, length of genal cones 0.03-0.05; length of rntennie
1.25-1.4; hcifii of uul tube 0.27-0.32. height of ~UMICICS 0.35-0.37.
Tenaife. Lu Merccdc~, 28-30 V 1947, 1 9. Gm Cpnuk: TiniuiS. S. BPrthdwis, 14 111
1950,19; ae iqúa, 28-29 111 199. i d; CNZ dc Tejeda 1450 m, 11-13 111 1950.1 9, 1
IV 1959.4 dd. inchidhg a holotype. 4 99; Suita B W, 31 111 1959,2 dd.
A.di&& sp.n cI& ruembkr the spceier desaihd bclow cQ.quifans sp.n.1. Hort pht
unknown
F b 104-112. ArYfUinüb diüm Loginova. sp.n.: 104 - hcad, from above below, 10s -
antenna. 106 - forewing of femalc. 107 - genitaih of femalc. in hteral vkw. 108 - geniw of
malc. in lateral view. 109- 110 - parameres, from insidc and above. 11 1 - apuc of pcnir, 112 -
e@.
o
!
I
Y
AryiamüiaequimsLoQnon, spnov.; fm 113-121.
Dcscribcd from 23 malu and femiler from Gran Canaria,
GreerryeiioV to mwtud yellow; b d dong antaior mugn of vutex. its foV- sclaites of
mctanotum and abdomen brown Patkm on muoaoúun in dder rpechnenr conrYtny of omp
bonds and rpotr Uppa putr of hud and thonx having dbtinct ügbtcdoumd uty whkh are
deiuer and loiyr thr in A&& q.n, rnd 2 pWr of the longcst setae w ged cona uurlly
light-cold inned dbrown
Foiavinl, Wrc thac of A.di&&, parhes oftcn ofcuniiy in ccll, of apial put, csll c da;
rninute upuficll rpim in uch eeU. in cell c fomiiry ovil rpot in middle.
Fe&.
plate b d y twke u h¡@
Donrl phk FnnUed ntar anur. fainüy concave abow (lateral vim). Triangular wntnl
u dorsal, with triangular bLder in middlc of interior mu&r embmbg
sidcr of dod phtc
Mak. Anal tubc *puLig ap"d. almort as high as psnmuu lod genital scgment. Param~rrr
iameüate. (heir nuroriy apicca turned fonvud and inwud their st~~tuie
ir vmy pcniiur. the
posterior andca of thai apiw be@ contiguwr (rim from above. Fig. 119). The structum of the
lpcxof thepn**hpecullr.
MuaimCatr Of tnnrle: la@ 267-3.12: kryh of fütCWing 225-2.55, ~idth 0.97-1.05;
width of hud 0.76471, width of vatex 0.45-0.49, ie~@th 0.15-0.17. length of wai cones
0.07; luigth of antemnc 1.42-1.6; lcngth of dorsai platc 0.7-0.75. Munuementa of mile: failth
255-3.0; k@ Of fOWiiU, 2.1-2.45. width 0.85-1.05; width of hed 0.78-0.8, ndth of
Vertu< 0.42-0.43. kyth 0.16-0.18; kryth Of pd COlKI 0.05-0.07; ICQth Of Mtainie
1.4-1.52;h&htüf ~aibuO.27-0.28.hd~toípuun~rtt0.3.
Gran Camia. Valb de TejmJi, 28-29 111 1949.1 9,2 dd(on C)Msus); Tirajana S Butholomc,
14 III 1950. 3 ¿¿, 1 9; CNZ de Tejeda, 1 IV 1959.1 d, 1 9; Santa B~igida, 31 111 1959.5 99;
Atairya, 26 VI, 4 dd. mClud@ a hdotype, 5 99 (kg. R. Fw).
Host plant unhom; colltcted once from Qtirur A.equifonr *.a. wu tonittimu rrcded
from the ymc gcogqhiul dbtrietr as A.di&lo sp.n; the two spceics are most probrbly rympatrlc.
llley are clorcly relatrd.

21
T
i
Fb 122-130. Arytainübpücohio Loghwa, sp.n.: 122 - hud. from r h e uid bdow, 123 -
anterI4 124 - forrwmg of femalc. 125 - gcnitrüi of fandq in hted vkw. 126 - pitrllrof
mak. in Ltenl view. 127-128 - pinmeier. fmm inde Ud .bovs. 129 - .pcx of p&
130 - CS.
’
Arytainilb pileobta Loginovp, sp.nov.; fi 122-130.
ücrribcd from 22 males and females from Tencrife.
Coiourcd iikc A.cquiianr sp.h, but in oider specimcns the veas with brown pigment are
snler. when yellow pitchcs occur on the forcwing. they are found oniy beside series of marginal
spiner Setic on thorax short and sparrr on uppcr prrt Forcwings oval, wkkst apierliy to middle,
erch ecU with minute superfxll spiner
Femilc. Do& piate concave ibove. no1 dhtcd anur Ventral platc triangulu, with a vcry süght
düaüon in middle of interna¡ mqin, ike that in A.dilufa sp.n.
Mak. A ~ r tube l nurow. like ihor of A.diluta and A.cquitanr spp.n., but bwcr thm
PUU~CKS uid genital rgmcnt. Parameres similar in form to those of A.hakoni Log., A.cognaia
Log. and A.innrba sp.n.; their anterior mugin hwing I mil trLnykr proas bililly to the
Iapering apcx and covd with hrge pecic; ipkd margin rmight, hebe of puunercr contiguair
at posterior angier of rpicei (vicw from ibove; Tic 128).
Muniiemaitr of femde: iength 2.5-2.87; length of fomwinga 2.05-237, width 0.87-1.02;
width of hud 0.7-0.76. width of vcrtex 0.41-0.49, kngth 0.15-0.18, iength of genal conea
0.07-0.1; iength of antennie 1.35-1.5; iength of dorui phte 0.65-0.67. Meuinmentc ofmak:
kngth 2.5-257; kngth of fonwmgs 2.02-2.25, width 0.85-0.95; width of herd 0.66-0.7, width
Of mteñ 0.4-0.43, hfih 0.15-0.17, kWh Of ~ nrl W(KI 0.07; kWh Of ~lte~uu~ I.45-1J;
height üf mal Nbe 0.25-0.27, he@t of P.IMIWW 0.22-0.25.
Tencrife: La Mcncdu, 1 9 (kg. R. Fny. N 1325). 28-30 V 1947,l ¿, 3 IV 1949,19,12
iV 1954. 7 ¿¿, inciuding i hoiotypc 5 99 (iq. R Remane); Orotiva, 3 d¿. 1 Q (*r. R Fny);
Bailadcro 700 m, 23 IV 1950,l ¿. 2 99 (on Lwrici).

In the SlNChire of udi put of the body, Apikobta ?.n. bcars a grcat rcsembluice lo two
representatives of the pnus from Morocco. A.cognara Lo& and A.hakoni Log, and 10 A.ineuba
sp.n from Mideir Thcr rpocies ue pnctidiy cioriy related. Unfortunatcly their host piant
rcmhs unknown, but thcir food source may be some kpminous ShNbL
ArifainUlo incuba Loginova, *.nov.; fi 131-140.
Dcsaibcd from 19 males and femalcs from Mdeira.
Only onc of the males ir grccn. the othcr Ipecimens are unifomly yeiiow without any pattern
on the thorax Ud abdanen. Antennae yellow or dirty yeiiow, up lo 3rd segmcnt, thc coiour then
grduiuy ch- apicad to dark brown. Fonwmp hyaiine. scldom faintiy whitish, no yeilow
patches in ecllr, vcins brown
Hcad U wide as tlrorax. dorsal sctac abscnt. Fonwings with anterior and posterior margins
more or ku po~Uel. mwiy roundcd at apial m8rgin with apcx in middlc of ceU n Minute
upeflcid Ipinrs in iu dlr Gcnai cones smiu ind WOW compucd with thorc of the othcr
qwcks and havhg the lrpe o€ obluraangicd tukmiiu.
Fe-. Dorsai piale rweUcd in anal rcgion, bu1 tapcriiy yduiuy. dody aimost straight.
Ventral piatc Me. b d y more than twiec u @e u dorsal. the wide biader on ik interior
margins not wekd u in Aquita~
but apprerrcd LO ddes of dorsal piatc (htenl VKW).
Maie. Anal tube wide. resembling that of A.cognata and A.liobni, more than twice as wide as
puuneres at ihcir base. lowu than pupmerer. Puuneres tapcring ipicad, with only fhrly defined
projstioní. not distendiry the interior marginK thek apicai mapins with acutc-andcd projections
birned fonvardr; thc hdLe in contact owing 10 contiguity of uppu parts of apiul myins (view
from abwe; Fb. 138).
Meas~mncnts of femaie: kngth 3.25-3.75; iength of forewings 2.75-3.1. width 1.07-1.2;
width of hud 0.77-0.87. width of vertcx 0.47-0.52. length 0.17-0.22, iength of gcnai cones
0.06-0.07; length of antennac 1.72-1.9; iength oí dorsai piate 0.75-0.85. Mcuunments of male:
iength 3.0-3.35; iength of forewiny 2.55-282. width 1.0-1.12; width of hcad 0.75-0.85; width
d vertcx 0.45-0.5. length 0.2-0.22, lcngth of pd cones 0.07; iength of antennae 1.77-1.85;
he@t Of uul tube 0.27-0.3, hciJit Of DuUnenS 0.35-0.37.
Madeira: Queimdas, 24-26 VI 1957,4 bd, including a hobtypc, 7 99;Ro&, 24 IV 1959,
3 óó, 2 99; Boaventura, 17 v 1959. i d. 2 99.
As alrcady mcntioncd. A.incubo sp.n. is vuy cioseiy reiatcd to A.cognuta Lq., A.hahni Log
and Apileobia sp.n.
Aryrainüb pmgnata Loginova ?.nov.; fis 141-142.
ücrribed frum a malc and 3 females from Gran Canaria.
üody iight yellow. vertex and oftcn pattern on meronotum orange; abdominal acierites
sometimes browtwpottcd. ventral piate and dorvl platc in puticulor builly brown Hed and
thoru wilh üght-coioured rctae abovc. hud U wide as thorpx. Ced cones comprntivdy @e.
Ud more than haif iength of vertu

.
31
Female. Dorsal pktc concave above. widcst in anal region. Ventral plate without dhtions on
convex interior marginr
Mile. AnJ tube mow. with apex somewhat bent backwards about 1.5 times u wide as
parameres at thcir bua. Parameres ükc hose of A.incuba sp.n., but more shatply tapering from
third of thcir height towardr apex, Iacking projections along anterior margin and denrely covercd
with long retae both anteriorly and porteriorly; lamellse of parameres contiguous almoot
throughout more or ICU stmight apical margin (vicw from abwe; fig. 142).
Mcasurements of femalc: Icngth 2.67-2.75; Icngth of forewings 2.25-2.3, width 0.92-0.95;
width of had 0.67-0.7, width of vertex 0.4-0.42, length 0.17-0.18. kngth of gnJ cones
0.1-0.12; length of antennae 1.15; lcngth of dorsal plate 0.7-0.75. Mcasurements of male: length
262; length of foming 2.15, width 0.82; width of head 0.67, width of vertex 0.4. length 0.17,
length of genal cones 0.1; length of antennae 1.25; height of anal tube 0.35. height ofparameres
0.27.
Gran Canuia: Valle de Tejedo. 28-29 III 1949, 1 d a holotypc, 2 99;
Tinjana. S Bartholorne,
14 III 1950, 1 9.
A.pugMta sp.n. hu thc grcatest reaunblancc toA.pi1eolota sp.n. Host plan1 unknown.
Arytainüh umbonata Loginon. sp.nov.; fig. 143-152.
ücscribed from 71 insrets from Madeh
Body grcm to pak yellow with oranp dorsal pattcrn and orangc abdomcnal scleritcs, often
bcaring blown spotr Forewing hyalinc. in ohicr insccts usually yellowish at apex; coloration
rtnngthcning towards apical mugin; veinr brown.
Hcad u wide u thorax, pnd cones very short, 1/3 or 1/4 length of vcrtcx. dhcted host
downwardr No dorsal setac. Fominga with anterior and posterior myins more or leu parallel.
widcst in apid Ihird, dightly cuning inwarda to Cu, along the apical myin. Minute aiperfictl
spincs not approaching veins, foming andl wrl spot in cell c.
Fcmaie. Dorsal plate düated in anai region. dightly concave above. Vmtnl pkte dutinguishcd
by long and narrow blades along interior myins and rtrongly diiated pprt in middle of lower
mugin (lateral view).
Malc. Anal lube strongly düated and bladc-üke postaiorly, widcst in fust third of hcight, higher
than thc parameres, and as high as genital rgment. Parameres rectuigulu (lateral view), thkk
posterior view. with dent interiorly in middk of anterior margin. Apex of parameres with long
widc projection. its widc bhdc being extended vutically. Parameres apicilly conIiguousat vertical
rnargins of iherc projections (dorsai and anterior view).
Measurements of female: lcngth 2.9-3.37; length of forewings 2.5-2.7. width 1.0-1.1; width
of hcad 0.71 0.77.width of vcrtex 0.43-0.47. Icngth O. 17-0.2. Icngth of pnal cones 0.05-0.07;
lcngth oí aniennae 1.5-1.75; Icngth of dorsal phte 0.6-0.75. Musurementr of mrle: lcngth
2.75-3.05; length of forewings 2.2-2.5. width 0.85-0.95; width of head 0.75-0.77, width of
vertcx 0.42-0.47, lengtb 0.17-0.2.len&th of genal cones 0.05-0.06; length of antennae 1.52-1.8;
height of anal Nbe 0.3, height of parameres 0.17.
Madcua: Sena de Aqua, 2 VI1 1957. 7 dd, 6 99, 16-17 Vil 1957, 4 dd, including a
holotype. 3 99, 20-21 IV 1959. 14 dd, 16 99(kg. Lindberg and Panelius. on Lyrminowe);
Curral das FreirasVvco Gil. 8 V 1959.7 dd, 13 99 0%. Lindberg and Paneliud; Ponta de Pargo,
23 IV 1959.19.
A.umbonaru sp.n. rcsernblcsA.incuba sp.n. in the sbucture of the fonwings, had and genilalia,
but ir clearly distinguishcd from it by the pcculk struchire of the ventral platc in thc female and
the parameres in thc male.
Aylia Ceoffry, 1762.
Thrcc spccies of ibis numerous genuí. two of them new, wcre discovmd in the Canary Idandr
Jp
Fin 143-152. Arytainilh umbonota Loginova, ip.n.: 143 - hud. from above and below, 144
- antauu; 145 - fmwing of fmirie. 146 - genitilir. of femdc, in lateral vh. 147 - ganitllk of
161 u
mde, in iaterai view, 148-149 - puunwes, from inside and above, 150 - apcx oí peak 151 -
hind 1%. 152 - egg.
Psy& arhntica Loginovr. ap.nw.; fw 153-160.
Duuibad fran a series of 184 inrtcts from Madeira and Tenerifc.
lñe insecu am p n at cmagencc, bu t m n change 10 yellow, then in yellowsrgangc-bnwq
abdomen latcnlly and genitlür sometimes crlnaon. Vertcx and dorsal pattun vivid o-. uid
mow stripes bemeen thc budi of pattern ivory-coloured; thonx ventrally md IatuiUy with
brown-bkk spots, legs yellow to brown. Forewingr tnnaiuccnt. colourleu to dirty ycllowkh. with
yellow band Nning along apid third of cüvus, ptcrostigrna brownish, minute superficial sphier
dark.

.
32 M. M. ~OWNOVA. F'quidrof rheCanq kluidraod
Hcad u wide as thorax. bent down through mgle of 40' or wcn more; gcnd cones u long as,
or more often lonpr thra vertex, rct not vuy vide aput. narrowly conical, rtrongly conave
extermlly. directed forwudr and downwardr Forrwingr widcrt in apKI1 thud, widcly rounded
aiong apiul margin, pterortigma loq, with dutinct R; minute superfiial piner in ueh ce4 leavins
spaee Jong veinr
Femaic GeNtllh cuneate. dorsal pbte di&hUy concave above.
GMUEMATMWES BKwa;icna 81 33
MJC A d tuk Sshapcd. aincd. not wide. plnmem Wrs thw ofhatbppínqu Log., but
with apvrl part morc disUncUy tapering and extended backwards rnd inwrrdr The puunau in
them rpc~Wr ~ V dLtinct C ~tlhier b W y rnd po~ariorfy (cC. Lodnov.. 1966: 22-23).
Mwurrmenta of fem.b: kqth 3.0-3.75; le@ of fomings 2.55-3.0, width 1.02-1.27;
width of hed 0.72-0.8, width of vertu 0.45-0.47, 1- 0.17-0.2, hgth üf ~CMI concg
0.2-0.25; kngth of antennac 1.07-1.25; length of dorsal pbte 0.75-0.9. Meuwmcab of mrh:
Iength 285-3.25; length of fomings 237-275, width 0.92-1.07; width of hud 0.66-0.74,
width of vertur 0.4-0.45, IenN 0.17-0.2,kn~th of gcd cones 0.2-0.22; height of uul (uk
0.25-0.27; hei@t oí p ~runf~~~ 0.17-0.22
M~deira: Pie0 Ruivo, 27-29 VI 1951,22 dd, 22 99 (onEricaarbm);Poim-A&ito. 19 VI
1957, 12 dd. 8 99 (on Lartus); QuQmdu, 24-26 VI 1957.1 9,14-16 V 1959,l d, 1 9 (m
StorA); Boanntun. 17 V 1959,4 dd, 8 99; Sem de
solir); Ribein Frb. 3 V, 1 d.5 %?(kg.
Aqua, 20-21 IV 1959.1 9; Ribclra Brava, 20 IV 1959.20 dbincludiq a hobiype. 29 %!ton
Prunur); So Vicente, 24 IV 1959, 1 9. Tende: Laa Mede 30 1 1949, 1 9.12 IV 1954.2
99 (ki. R. Rcmw); V i r 1500 m. 27 11 1 9 5 O d f r u 1300 m. 19 V 1950. 1 9:
Valle de S Andri4 2 III 1950. 1 d; GAdilk, 26 lI~i95& 1 d; Bm.dcL Inf&, 24 11 1950,ZO
66,21 YY (m M&).
In tbe complex of wUow-fe#tiiy spcckr of the pnus Psyk thb wier ir the ninth
rcpnlcntatlve of thc group ftwphih (Loginovr 1967). It mort dosly rsrmbka Rpn~pinq~~
SchuL and Rrubpropinqua Log. DUtinctive fatures in the stNcturc of the hud, forcwinp iiid
gcniwL rnd its irobted ara indicate tlutP.ut&ntica sp.n. ir an indepcndent rpccicr Its hwt phnt
u clurly thc wiuow.
PIylh exima Loginova, ?.nov.; fig~. 161-169.
F& 1531-160. Psyih atlontica Lo&ova, íp.n.: 153 - hud. Crom rbwe and below, 154 -
forewiy of mile. 155 - genitllk of femde. in bieril view. 156 - gcnitslll of mak in hterd view,
157-158 - puamcres, from insidc and above. 159 - apex of penis. 160 - hind leg.
' Ducnbcd from 29 males and fanaiu from Tencrife, 26 of thm cobcted by Dr. R. Remane in
1954.
Somc specimcnr pm-onng with a brown abdomen, thc mapdty much darkrcolw~. hcd
Ud thm omgc with a brown pittm above and rpotr on rider Ud bclow. Vutex iloq
cpiuullil suture. pmnotum. prvrcutal uyicr of mesonotum üght yeüow. pnil mnu yeüow to
omge Antennae to 5-6th regnuit duty ydlow, regmentr 3-6 ipicdly brovn. mae rpieri
segments brown bkk. Le@ dirty yeUow with brovn lana Forewinga transpucnt. ydowish.
veinr, minute superficial pincs and apid hrlf of drvur on both pain of wings brown Ventnl
rleriter of abdomen browmblrck, the hterqmentd mcmbruie reddish 01 onnge. uul tuk rnd
parameres of mile dirty yellow.
Hud U Wide UiOnX. StrOngIy bent f0Wudr (thr~gh Of 30'). g C d cones dincted
vertiully dowmds, almort u loq as vertex. sct wide 8pUt. nurowly conM artUidy
roncavc Antuuue rlmost 1.5 tima u loiy u had Forewing widert in ipkd third thm waüy
munded. ptemstignr wide. b@y almort u wide u adjoining rcgion of ceU r, tetqimlosing at
level of bifurution of M. minute superficial rpiaer dcnscly oct. but not clore lo veinr (4162).
Fermlc. Both platcr tapcring rpiud. donrl pira EOIL~.YC rbovc
Milc. A d tube convex ~tuiorly, muvc postaiorly, u high as parmeres, but lower thm
the hemkphedd genitll regment Paramera u narrow Lunellre with a nun>wed ipur. dnwn
fonvrrds (lateral view and view fmm above), thkk posterior view. ilong posterior mugin curyinl
a comb of 1-2 liner of very rtrons brown pbteller Apiul put of penis having d&ht oval dilrtion.
ductur long.
Mcuuranents of femak: kwth 265-3.0; kqth of forewbgr 2.27-2.65, width 0.95-1.0;
width of he& 0.65-0.67, width of Vatu( 0.4-0.42. l~ngth 0.17-0.2, kngth of genal CWI
0.17-0.2; length of antcnnae 0.97-1.07; length of donil pbtc 0.62-0.75. Measuranentr ofmile:
kngth 2.45-2.67. kngth of forewings 2.07-2.17. width 0.85-0.9; width Of hed 0.65; widfhof
vemx 0.37. length 0.15-0.17. kngth of gcd cones 0.15-0.17; length of antennre 0.92-1.05;
height of a d tube and paramcru 0.25.

.
34
35
T~za Fhter, 1848.
7iioza unlcoc 4) - Yadeira, PnL Fonnou pr. FuncW 28 IV 1959,19 d9.
On Urda dioicn, U urew u S Antonio, 21 V 1947, 5 d9.4 II 1949. 5 d9 (onLmuur);-
Aguim, 17 11 1949.17 ¿%Puerto de la CNZ, 2-4 11 1949,l 9; Fuente fria 1300 m, 19 IV 1950,
mVueita de Tyuiw 700 m, 16-17 11 1950. 3 99. Gran Canaria: Moya, 14 d9 (iet R
Std);CmzIle ieJw& 1 rP 1949,2 dd.
On Laumr nobilk. Lauur canaden* the nymphr cause malformationr in the fouYq
putfflkrly in young shootr; the marginr of the luvu curl downwudq forming gaüs, in whkh the
nymphs develop. Therr are hvo pnuationr a ycpi íin the Crima, the Caucuur wíthin the borda
of the USSR), and the adult inaectr hibematc. Widely dirtlibutcd throughout the Medturioun
corsts of Europe up to tbe Cauaniq reportcd fmm Algcria. Known in mon northun reghnr up to
En- and Swcden, but only in hot-houset Importcd to North and South Americo, A put.
Figs 161-169. Prylh uima Loginova, sp.n.: 161 - hud. from abovc and bclow. 162 .-
forcwirq of femalc. 163 - genitJir of female. in lateral vkw. 164 - donrlphte. from abs); Si- 1211 1949.1 ¿;PII&O de S JUM. 23 11
1950.6 d9; Bur. del Infierno, 24 U 1950,12 dQ; Tunaimo 600 m, 22 11 1950.1 9; Coja-Tejina
500 m. 23 11 1950. 1 9; aipn Adcje Querta de la Puor 1200 m. 25 11 1950, 1 ¿; ciwlu
2300-2500 m. 28-30 111 1954. 1 9 0% Rmm). Cna Cuwi.. Lu Phw 5-6 VI 1947,4
d9 (on Cñenopudium); Condo. 9 111 1963. 5 d9; Gddu. 23 U 1949. 10 d9; Mrrpdomy
24-26 11 1949,7 d9; Suih B&i& 21 U 1949.1 9; ANCU, 2 IU 1949.1 d. LMZUO~CH~~~,
10-15 111 1963, 14 69; Fms 18 111 1963, 8 d9; P d u ~
del Qiacha, 13,22 111 1963,l 9.
Fuuteventurr, Chikgua, 4-14 111 1949,l d.
On onnuai rpcier of Aorpu and Climopodium Centnl and Southem Europe, Klukhstui.
Middk Asia of thc USSR. Morocw.
On his numerous travels, Profesor Hlkon Lindberg, m outstanding IutUdkt and
man of science, cnthuriasticaiiy coiiseted reprerntatives of various inrct goupr His
collections of psyliiis from Morocco, thc Canuy . Isiands and Madein wbn
undoubtedly not intended to give a complete picturc of the fauna of thh lpoup in
thor areas, being evidentiy more or les incidentil. Nevythcles, they are of a p at
significance as the first matcriai offeriq infonnation on this intensting BIOUP in the
extreme south-mst of the Palaearctic. The collections from the Canaries and Mpdeira

.
are especirlly valuable. since they give some conccption of ihe distinctive character
and weaith of the fauna of thoa isiands.
Rylloideu are phytophagou wcking ihcects of s d &.e. They show very narrow
trophic spccidization, being restricted to ccrtain pht~, SO and their divertity and
genetic differentiation are thus primarüy detcrmined by the -ter of the pnmt
vegetation and its genesir in a @ven territory.
Macaronesia ir an independent floral Mon closely COMCCted with the Meditemaman
district and the Palaeotropics and having some boreal and neotropical elements in
its flora (Iijinsky, 1937). The Canaries. at the eastem liMt of the *on, lie in the
zone with an arid climate and semidesert psunmophytous wgetation, almost wholly
devoid of arboreous species. The Westem Canariea are mountahous and have a
luxuriant vegetation. Altogether, 1300 spccies of vwñllu @ts grow on the Canaries,
806 of which are aboriginal, haif endemic md 490 imported.
In the lower zone (up to 700-800 m above sea-levei) the vegetation has a
subtropical-African character. Four vertical zones can be tnced in the mountainous
region - the laurel forests at a height of 700-800 m are gradually nplpced by the
maquis; at a height of 1600-2100 m are forests ofpimcs cana&nsis, and at a height of
2000-32000 m the dopes are covered with scnibby uphd derrts, with bushes of
Spartocytisus supmbiur
Among the most common laurel spccies are Persea úuika, Lautus crmorirnuí and
Ocoreu faiens; the maquis eonrut mortiy of arboreous heathen, su& u Ericu
uhomu, of üex. cümbing Convolvulus cmuviensir, and ieguminous shnibs, e.& Cyricus
pmli/cnrs and Adenocurpus viscosur
Iñc climate in Madeira ir more equabie than that of the Cwies, though it is
ninier and the average annual temperature ir somewhat lower. There are no pine
foreats and mountainous deserts. A zonc with ihra distiishable phytocoenoses
foUowr the zone of laurel forests: there grow ccrellr, upiand shrubs. including
Vpcemium mudemse, and hcaiths with Erica arboreu and other phts.
Thirty four pcr ant of the flon of Madeira is endemic; thesc endemic specier are
cither relict spacies from Madeúa or nlated to South European species. nie phts
growing on open land in both Madein and-the Canaries are dortly endemic.
Considention of the zoopopphical composition of the North Macaronesian
isiands is rertiicted by the meagreneu of the information avrilable on the food,
biology and distribution of these.
Ea& of the 29 species treated here are reported from these isiands for the first
time, threc species have recently been described by the author from Morocco and
seven are more or leu widely distributed in Europe and Asia.
The most typid feature of the fauna is its Palaearctic - Mediterranean chancter:
the gnat majority of the spccies (24-25) are representatives of the humid subtropical
Mediterranean region. Thiu U not surprishg, since the hbtoriul connections and
preant procimity of the idands to the coasts of Europe and Africa have resuited in
great similarlities betwan thcir climate, vegetation and fauna.
Howcver, of the 20 new speciea, Psyl& atiantica sp.n. aionc differs iittie from its
cloaest Europan-Asian relations, i.e. the species of the group Itheophila, and
Aryfainil& pileohta and A.innrba spp.n. are geneticaily closely connectcd with two
I
species from Morocco. A . h i and A.cogmta me rcmahng 17 speeies are
strüungiy difiercnt. and the rmjority teem to be endgnic to Uie Cuurics and Mide¡¡.
Thur, the psylüd fauna of the irluidr ir mnrrkible for its diatinctive diuicter and
lye numbcr of endcmic specks and genen.
Am~ng the 10 spccier known from the Paiaearctic, Tdom urricoc ir a tnnrp.lrearctic
forest speciu. and Triozu obliqu U llro widely diatributed in that zoogeognphical
region. but OCEUR mortly in the uid mnc. Thra rpocks osighate from the uid
zone, Rhodochlanis puw@uncturur. DiaphonnO conlinuri and Co@osceniu viridis and
until now were biown oniy from Southem Moroca.
Two of thepe thm specier are reported from the deserts of the eartem Uiuids
LPNIrote and Fuerteventure, while Cvirkiis U alro reported from Cmn Cinuir md
from Portugi. Thc npnsentativ« of the gcnus RhodochhnU are xeported from
annual dtwortp of the Owno- which are haiophytous and often occu
beyond the zone of deserta and stcpper. nie host pht for R puwipuncrurus ir not
yet known, the speeier is recorded from Morocco and Algeria.
The remaining fiw rpccics are of Meditemnean origjn, their faod sourcu are
typicai subtropicai arborsJ piants (Oleu. purcicis Lowus etc); like most of the new
species, thcy are widely distributed in South Europe and North Africa, and mm of
thmi penetrate far into the arid subtropics.
The genetic composition of the Ayl&ideu fauna from the Canary Isluids and
Madeira ir thu fairiy homogtneous, the grut majority of the species being of
subtropiul origin. There are no boreai speciea; the fact that 10 !% of the species
originate from the arid zone indicates a conncction with the Sachara depert fauna. niiS
connection ir les marked on the ihds Ytuated fpither from the coa of South
Morocco.
Litmhim
Copcner A.L 1970. Southern Afhcm Pguidic (Homoptcra). 1 - J.entSoc.rth.Af~, vol 33. No. 2:
195-200.
Hesbpffmhon C. 1951: Thc Alytiinini of thc SubhUy PIYlliay HaniptcrrHanop~
FmJV P~uid.e - Allh M& N8t Hh. SCK 12. ~014:417-454.
HedopHoniron C. 1961: Hempfui (Homoptcn): Psywdra - South Afr.Anim.Lüe, voL8.
uppSal8: 487-532
IlíjimkyA.P. 1937: Vegetition of the world. Aad.ScL USSR. Morccnw-hW 252-270.
Loginovu M.M. 1966: New rnd littiekoown psyUidr (Homoptsn) frm the USSR - Tniuctbiu
of Zooiogid Inbiutc of Acadany of Scknce USSR. 37: 3-35.
~ @ O W M.M. 1967: Renw on pd8cuctic prYuidr of íhe enur h~ui Cwífr. (Homopt~,
Plylloidu), uiaciitcd wiIh willowr - AnnZoolqici, 34, No. 7, Wunw: 427-461.
Lw'novu MM. 1972: On the fauna of Pglloidu (Homoptera) fmn Moroeco. - CommenLBioL.
47. H~lPnki-Hellln~~~~: 3-31.