28.12.2013 Views

j - Dipartimento di Matematica - Politecnico di Torino

j - Dipartimento di Matematica - Politecnico di Torino

j - Dipartimento di Matematica - Politecnico di Torino

SHOW MORE
SHOW LESS

You also want an ePaper? Increase the reach of your titles

YUMPU automatically turns print PDFs into web optimized ePapers that Google loves.

hard copy ISSN 1974-3041<br />

on-line ISSN 1974-305X<br />

La <strong>Matematica</strong><br />

e le sue Applicazioni<br />

n. 11, 2010<br />

A mathematical procedure for the evolution of future<br />

landscape scenarios<br />

F. Gobattoni, G. Lauro, A. Leone, R. Monaco, R. Pelorosso<br />

Quaderni del<br />

<strong>Dipartimento</strong> <strong>di</strong> <strong>Matematica</strong><br />

<strong>Politecnico</strong> <strong>di</strong> <strong>Torino</strong><br />

Corso Duca degli Abruzzi, 24 – 10129 <strong>Torino</strong> – Italia


E<strong>di</strong>zioni C.L.U.T. - <strong>Torino</strong><br />

Corso Duca degli Abruzzi, 24<br />

10129 <strong>Torino</strong><br />

Tel. 011 564 79 80 - Fax. 011 54 21 92<br />

La <strong>Matematica</strong> e le sue Applicazioni<br />

hard copy ISSN 1974-3041<br />

on-line ISSN 1974-305X<br />

Direttore: Clau<strong>di</strong>o Canuto<br />

Comitato e<strong>di</strong>toriale: N. Bellomo, C. Canuto, G. Casnati, M. Gasparini, R. Monaco,<br />

G. Monegato, L. Pandolfi, G. Pistone, S. Salamon, E. Serra, A. Tabacco<br />

Esemplare fuori commercio<br />

accettato nel mese <strong>di</strong> Ottobre 2010


A mathematical procedure for the evolution of future landscape scenarios.<br />

Authors: F. Gobattoni 1 , G. Lauro 2 , A. Leone 1 , R. Monaco 3 , R. Pelorosso 1<br />

1 Department of Environment and Forestry, University of Tuscia, Viterbo (Italy)<br />

2 Architecture Faculty, Second University of Naples, 81031 Aversa, (Italy)<br />

3 Department of Mathematics, <strong>Politecnico</strong> <strong>di</strong> <strong>Torino</strong>, <strong>Torino</strong> (Italy)<br />

Correspon<strong>di</strong>ng author: f.gobattoni@unitus.it<br />

Introduction<br />

Since the European Landscape Convention aims to encourage all European countries to define their<br />

landscape quality objectives, a new frame of mind in management and planning of territory is<br />

nowadays required as a point of reference for territorial government, entities and authorities, so as<br />

to advance towards conservation and protection of landscapes provi<strong>di</strong>ng positive impact on the<br />

quality of life of population. Moreover, planning efficiently at the landscape scale needs a crosssector<br />

cooperation and people involvement with planners and communities working closely in<br />

achieving positive landscape change understan<strong>di</strong>ng and in characterizing the relations between<br />

nature and society, which will integrate landscapes and their dynamics. (C. Petit et al., 2008).<br />

The methodologies for successful landscape and environmental planning have been severely<br />

challenged when classical concepts and models such as economic and socioeconomic development,<br />

ecosystems preservation and sustainability have been questioned (J. E. Vermaat et al., 2005). The<br />

actual challenge is to build transparent and flexible decision-making tools to be used in<br />

environmental planning and to embrace a broad range of stakeholders needs together with<br />

landscape management requirements.<br />

Landscape-scale geographical information provides an ideal framework for developing spatial<br />

strategies and to implement simulation models to support landscape planning identifying<br />

opportunities for change and anticipating and comparing the results of planning decisions. Model<br />

outputs can be used to elaborate thematic maps and visualizations of landscape scenarios as an<br />

effective and <strong>di</strong>rect way of communicating results and opportunities.<br />

In this framework, landscapes can be defined as spatially extended heterogenous complex systems<br />

organized hierarchically into structural arrangements determined by nonlinear interactions among<br />

their components through flows of energy and materials (we shall use the overall term "bio-energy"<br />

as introduced by Ingegnoli, 2002 ). Human activity has, also, strongly mo<strong>di</strong>fied recent landscape by<br />

means of land use and land cover changes and consuming of natural resources (Pelorosso et al.,<br />

2009). Alterations of natural equilibriums has pointed out several consequences on landscape<br />

capacity to furnish goods and service (Willemen et al, 2008) as bio<strong>di</strong>versity conservation<br />

(Tscharntke et al., 2005) and regulation of water regimes (Lindborg et al., 2008).<br />

Environmental available energy has been pointed out as a key factor in the explanation of many<br />

ecological processes and theories (metabolic and species energy theory, bio<strong>di</strong>versity conservation)<br />

recurring to <strong>di</strong>fferent indeces (e.g. Net Primary Productivity, NNP, Actual Evapotraspiration, AET,<br />

Potential Evapotraspiration, PET) (Hurlbert and Jetz, 2010; Currie, 1991; Carrara et al. 2010;<br />

Brown at al., 2004). A more complete measure of available energy, so-called Biological Territorial<br />

Capacity (BTC), was proposed by Ingegnoli (2002) considering a synthetic function of vegetation<br />

metabolism. As the importance of energy exchange, although it is rare for a landscape to be in any<br />

form of equilibrium and the landscape equilibrium concept is not yet clear (Perry, 2002; Turner et<br />

al., 1993; Bracken and Wainwright, 2006), it’s interesting to focus on which hypothetical energetic<br />

equilibrium state is going to be realized and the effects of human decisions on that equilibrium.<br />

Most of mathematical models are strictly related to air <strong>di</strong>spersion, hydrology and hydrodynamics,<br />

water quality, ground water quality, erosion and se<strong>di</strong>mentation, and so on, just taking into account<br />

each aspect of the environmental system separately and without looking <strong>di</strong>rectly at landscape as a


unique system and without understan<strong>di</strong>ng its intrinsic evolution mechanisms. All decision making<br />

involves an implicit (if not explicit) use of models, since the decision maker invariably has a causal<br />

relationship in mind when he makes a decision. So mathematical models able to explain the<br />

landscape evolution and to compare the effects of future scenarios on its evolution and equilibrium<br />

in time, are really needed not also to understand environmental system mechanism and behaviour<br />

but above all to better plan strategies for natural resources conservation management and landscape<br />

preservation.<br />

The environment is here considered as composed by several Landscape Units (LU) delimited by<br />

natural and/or anthropic barriers. An integrated GIS (Geographic Information System)-based<br />

approach was developed (G. Lauro, R. Monaco, 2008) combining an ecological graph model for the<br />

analysis of the relationship between spatial pattern and ecological fluxes and a mathematical model,<br />

based on a system of two nonlinear <strong>di</strong>fferential equations. These equations are mainly based on a<br />

balance law between a logistic growth of bio-energy and its reduction due to limiting factors<br />

coming from environmental constraints. The energy exchange among them will be more or less<br />

strong depen<strong>di</strong>ng on the degree of permeability of the barriers which can obstruct the energy<br />

passage from each LU to the other.<br />

A GIS-based mathematical model, based on the ecological graph and on the cited two <strong>di</strong>fferential<br />

equations, is presented and <strong>di</strong>scussed here. A study case in Central Italy is introduced and <strong>di</strong>scussed<br />

here just to show the importance of a such mathematical procedure to figure out planning strategies<br />

and environmental conservation and management actions with the support of rigorous methods able<br />

to fix and compare the effects of anthropic decisions on landscape.<br />

Study area description<br />

The study area (Fig. 1) for the model development, is Traponzo River catchment; Traponzo River is<br />

located in the northern part of Lazio Region and its watershed covers part of nine municipalities,<br />

lying completely in the Province of Viterbo.<br />

This stream originates in Monti Cimini relief and flows into Marta river so that Traponzo<br />

catchment, as a sub-basin of Marta river, has a total area of 475 Km 2 , with a mean elevation of 526<br />

m a.s.l. and a maximum of 979 m a.s.l.. The climate of this area is quite Me<strong>di</strong>terranean with a mean<br />

annual temperature of about 15°C and a mean annual precipitation of about 970 mm. The total area<br />

covered by urban sprawl is about 2.23 km 2 that is about 10% of the total urban area.<br />

Land cover class Area (km 2 ) %<br />

Urban 21.02 4.4<br />

Forest 126.36 26.6<br />

Non irrigated crops 203.04 42.8<br />

Pasture 22.01 4.6<br />

Orchards 88.17 18.6<br />

Irrigated crops 2.49 0.5<br />

Hedges 11.77 2.5<br />

Water bo<strong>di</strong>es 0.02 0<br />

Total area 474.89 100<br />

Table 1.


Fig.1. Study area


Model description and methodologies<br />

Gathering all the available information on the climatic, topographic, land cover, geological and<br />

pedological characteristics describing our study area, a GIS dataset has been created not only to<br />

represent the watershed but also to identify and derive the LU structure. Taking into account urban<br />

areas <strong>di</strong>stribution, primary and secondary roads network, soil features and elevation and weighting<br />

them through the application of Saaty matrix, a standard heuristic methodology has been fixed to<br />

split the study area into well defined LU. A number of 46 LU has been derived for this study area<br />

and an ecological graph for representing the energy flows between them has been elaborated taking<br />

into account the Biological Territorial Capacity (BTC) proposed by Ingegnoli (2002). The<br />

ecological graph, at first introduced by Fabbri (2003), has been applied here to quantify and<br />

correlate the energy production from a landscape unit as a flow of energy to its neighbours<br />

accor<strong>di</strong>ng to the permeability of the boundaries.<br />

The study area GIS dataset also allow to set up all the parameters needed to run the mathematical<br />

model structured as follows.<br />

It consists in two or<strong>di</strong>nary <strong>di</strong>fferential equations, of evolution in time, of the following two state<br />

variables, M(t) and V(t) :<br />

1) M(t), the mean value of the biological energy of the whole system, t being the time, given by<br />

m<br />

1<br />

M ( t)<br />

= ∑ M j<br />

, M j = (1 + Kj)Bj (1), (2)<br />

m<br />

j=<br />

1<br />

where Bj is an index measuring the BTC of the j LU, that is the magnitude representing the energy<br />

(Mcal/m 2 /year) that system needs to <strong>di</strong>ssipate in order to maintain its equilibrium state and its<br />

organizational level, and where the parameter Kj depends on several properties related to unit’s<br />

composition. In particular, in this paper we shall consider that it depends on 5 features and it is<br />

given by<br />

F P D C E<br />

K = ( K + K + K + K + K ) / 5<br />

(3)<br />

j<br />

where<br />

j<br />

j<br />

j<br />

j<br />

j<br />

F<br />

K<br />

j<br />

takes into account the shape of the patch borders,<br />

P<br />

K<br />

j<br />

their permeability,<br />

D<br />

K<br />

j<br />

the bio-<br />

C<br />

E<br />

<strong>di</strong>versity of the LU. The last two parameters K<br />

j<br />

and K<br />

j<br />

are related, respectively, to the relative<br />

humi<strong>di</strong>ty of the soil and to the sun exposition of the LU, and are introduced for the first time in this<br />

paper. We recall that all these parameters are defined in such a way that K ∈ [0; 1].<br />

For the computation of the parameters<br />

Conversely<br />

K<br />

C<br />

j<br />

w1<br />

A<br />

=<br />

h<br />

j<br />

C<br />

K<br />

j<br />

and<br />

+ w<br />

A<br />

j<br />

2<br />

A<br />

s<br />

j<br />

F<br />

K<br />

j<br />

,<br />

P<br />

K<br />

j<br />

and<br />

E<br />

K<br />

j<br />

are defined by the following formulae:<br />

, K<br />

where the w k are suitable weights,<br />

E<br />

j<br />

SWE W<br />

3<br />

Aj<br />

+ w4<br />

Aj<br />

+<br />

j<br />

D<br />

K<br />

j<br />

, see Finotto et al. (2010).<br />

NE<br />

w<br />

w5<br />

Aj<br />

= (4), (5)<br />

A<br />

Aj<br />

is the total area of the j LU, and<br />

j<br />

h s<br />

A<br />

j<br />

, A<br />

j<br />

,<br />

SWE<br />

A<br />

j<br />

,<br />

W<br />

A<br />

j<br />

,<br />

are the fractions of soil surfaces characterized by humi<strong>di</strong>ty, sub-humi<strong>di</strong>ty, south-west-east, west and<br />

north-west exposition.<br />

Moreover:<br />

M = , B max { B }<br />

max<br />

2B max<br />

max<br />

= (6), (7)<br />

j=<br />

1,...., m<br />

j<br />

NE<br />

A<br />

j


in other words M max is the maximum value of biological energy that the environment under<br />

consideration can exhibit accor<strong>di</strong>ng to the maximum value of bio-potentiality expressed by the m<br />

<strong>di</strong>fferent LU. These last definitions lead to fix the bio-potentiality index as<br />

B<br />

b T<br />

= (8)<br />

B max<br />

which, thus, turns out to be confined in the range [0; 1].<br />

2) V (t), the fraction of the total territory’s surface occupied by areas characterized by high values<br />

of Bj ( high values correspond, for example, to wooden areas).<br />

The proposed equations read :<br />

' ⎡ M ( t)<br />

⎤<br />

M ( t)<br />

= cM ( t)<br />

⎢1<br />

− − k[ 1 −V<br />

( t)<br />

] M ( t),<br />

M<br />

⎥<br />

(9)<br />

⎣ max ⎦<br />

[ 1 −V<br />

( t)<br />

] h U V ( ),<br />

V ' ( t)<br />

= bTV<br />

( t)<br />

−<br />

0 0<br />

t<br />

(10)<br />

They read as a balance law between a logistic growth of bio-energy M (t) and its reduction due to<br />

limiting factors coming from environmental constraints expressed by the coefficients c, k, b T , h,<br />

Uo.<br />

Having already given the definition of b T , we now proceed to deal with the other parameters.<br />

c∈ [0; 1] is the connectivity index which is a function of the energy fluxes F ij between each pair of<br />

confining LU, namely<br />

S S<br />

Mi<br />

+ M<br />

j<br />

Fij<br />

=<br />

2<br />

Lij<br />

⋅ ⋅ p<br />

P + P<br />

i<br />

j<br />

(11)<br />

L ij being the length of the boundary between the i and j LU; P i and P j being their perimeters, while<br />

p is the permeability index of such a boundary. Moreover k is the ratio between the sum of the<br />

perimeters of the impermeable barriers and the perimeter of the whole environment. h is another<br />

environment impact parameter defined as the ratio between the sum of the e<strong>di</strong>fied areas perimeter<br />

(both compact and spread) and the total perimeter P of the whole environment. Finally U 0 ∈ [0; 1]<br />

is the ratio between the surface of the e<strong>di</strong>fied areas and the total area S of the system. U 0 will be<br />

computed as the weighted average:<br />

U<br />

0<br />

w6U<br />

c<br />

+ w7U<br />

s<br />

= (12)<br />

S<br />

where U c and U s are the surface fractions of compact and spread e<strong>di</strong>fied areas, and w 6 , w 7 suitable<br />

weights. Note that conversely to the other parameters of the model, k and h may assume values<br />

greater than one (Finotto et al., 2010).<br />

In order to get a detailed evolution of the environment, one can integrate the equations (9-10) from<br />

suitable initial data M(0) and V(0) which can be recovered by the ecological graph. The model<br />

provide these scenarios correspon<strong>di</strong>ng to the following four equilibrium solutions of the <strong>di</strong>fferential<br />

equation system:


(1)<br />

(1)<br />

(a) M = 0 ; V = 0<br />

(2)<br />

(b) M = 0 ;<br />

( 2) bT<br />

− hU<br />

0<br />

V<br />

=<br />

b<br />

T<br />

(c)<br />

(d)<br />

M<br />

M<br />

M<br />

max<br />

( c k)<br />

(3)<br />

= , V = 0<br />

c<br />

( 3) −<br />

(4)<br />

=<br />

M<br />

max<br />

[ c − k( 1 −V<br />

)]<br />

c<br />

e<br />

,<br />

b<br />

=<br />

hU<br />

b<br />

( 4) T<br />

−<br />

0<br />

V<br />

T<br />

As it can be easily understood the first equilibrium is quite negative since it prevents an<br />

environment where production and <strong>di</strong>ffusion of biological energy is negligible and no areas of high<br />

ecological quality are present. The second scenario is that of a territory strongly fragmented where<br />

<strong>di</strong>ffusion of biological energy between the LU is again negligible but some area of high quality<br />

vegetation is still present. The third equilibrium corresponds to a territory characterized by some<br />

production and <strong>di</strong>ffusion of bio-energy but low vegetation quality. Finally the last scenario is that<br />

more favourable since strong production and <strong>di</strong>ffusion of biological energy between the LU allows<br />

to guarantee an ecological settlement of high level of bio-potentiality. The stability analysis (Finotto<br />

et al., 2010) has shown that the above equilibrium solutions can be obtained, respectively, when the<br />

model parameters satisfy the following inequalities:<br />

c < k and b T<br />

< hU<br />

0<br />

(I)<br />

khU<br />

0<br />

> cb T<br />

and b T<br />

> hU<br />

0<br />

(II)<br />

c > k and b T<br />

< hU<br />

0<br />

(III)<br />

khU<br />

0<br />

> cb T<br />

and b T<br />

> hU<br />

0<br />

(IV)<br />

The ecological graph representation together with the solutions of the mathematical procedure can<br />

be derived through a NetLogo model application that give us an easy tool to model the equilibrium<br />

states of landscape evolution starting from the initial, actual con<strong>di</strong>tions as described by the<br />

ecological graph itself (Gobattoni F. et al., 2010).<br />

Results and <strong>di</strong>scussion<br />

As an assessment of the ecological behaviour for an environment, the ecological graph assigns a<br />

node <strong>di</strong>mension proportional to the available energy and a link between LU with a <strong>di</strong>mension<br />

proportional to the flux of energy. The energy exchange among them will be more or less strong<br />

depen<strong>di</strong>ng on the degree of permeability of the barriers which can obstruct the energy passage from<br />

each LU to the other.


Fig. 2 Ecological graph representation for the study area.


Applying the proposed LU identification methodology and deriving all the required parameters<br />

needed to set up the mathematical model, this summarizing table can be obtained:<br />

Total area (km 2 ) 475<br />

High bio-potentiality area (km 2 ) 124<br />

V 0 0.26<br />

b T 0.1606<br />

M max (Mcal/m 2 /year) 2.3 E+08<br />

B max (Mcal/m 2 /year) 1.5 E+08<br />

U 0 0.02<br />

h 1.96<br />

k 0.75<br />

c 0.0374<br />

V e 0.72<br />

M e 0<br />

Table 2.<br />

The values in Table 2, have been entered in NetLogo model elaborating the proposed <strong>di</strong>fferential<br />

system. In particular, b<br />

T<br />

, h, U<br />

0<br />

, M max , k and c seems to satisfy the inequality (II) so that the<br />

equilibrium correspon<strong>di</strong>ng to he second scenario turns out to be stable and the system evolves<br />

asymptotically to this equilibrium state. The system tends to keep areas with high value of biopotentiality<br />

but they are isolated in landscape pattern and the fluxes of bio-energy between them<br />

seem to be limited. As a consequence, the dynamic evolution of this environmental system shows a<br />

trend toward a good production of bio-energy but with a limited <strong>di</strong>ffusion of it, and, then, with<br />

limited fluxes between LU. A low connectivity index (Table 2), <strong>di</strong>scloses to this equilibrium state<br />

as a solution of the <strong>di</strong>fferential equations system, since the lack of connectivity preju<strong>di</strong>ces the<br />

energy exchange between LU even containing high BTC values areas. The landscape fragmentation<br />

provoked by a large urban sprawl phenomenon and by the rich and structured roads network is<br />

reflected on the confined and obstructed energy fluxes between LU. The need of opportune<br />

planning strategies and actions to reduce fragmentation favouring the energy fluxes between<br />

ecosystems and preserving bio<strong>di</strong>versity, has to be underlined since the actual landscape pattern for<br />

the study area shows a low response in terms of connectivity and energy fluxes.<br />

Conclusions<br />

If the ecological graph is a powerful tool to represent connections efficiency between LU and to<br />

identify high ecological values areas to be protected and compromised ones limiting energy fluxes,<br />

on the other side a mathematical model evaluating the potential effectiveness of natural resources<br />

on the long term, is essential to assess the <strong>di</strong>achronic evolution of landscape as a unique system.<br />

An integrated approach combining the ecological graph and the mathematical model, allows to face<br />

the great challenge of planning and management under sustainable environmental and economical<br />

con<strong>di</strong>tions since it can represent a powerful decision system support to compare effects and impacts<br />

of alternative scenarios and actions (evaluation of new roads and urban development plans).<br />

Not only for a description of the available energy at LU scale but also as a tool for evaluating the<br />

equilibrium trend in landscape evolution, this mathematical and GIS interfaced method can help in<br />

understan<strong>di</strong>ng environment response and dynamic change in time to correctly manage and preserve<br />

natural resources and ecosystems.


References<br />

Bracken L. J and Wainwright J. 2006. Geomorphological equilibrium: myth and metaphor? Trans<br />

Inst Br Geogr, 31:167–178.<br />

Brown J. H., Gillooly J. F., Allen A. P., Savage V. M., West G. B. 2004. Toward a Metabolic<br />

Theory of Ecology. Ecology, 85(4):1771-1789.<br />

Carrara R. and Vàzquez D. P. 2010. The species energy theory: a role for energy variability.<br />

Ecography 000: 000000 doi: 10.1111/j.1600-0587.2009.05756.x.<br />

Currie D. J. 1991. Energy and large-scale patterns of Animal- and Plant-Species Richness. The<br />

American Naturalist, 137:27-49.<br />

Fabbri P., Paesaggio, Pianificazione, Sostenibilità, Alinea E<strong>di</strong>trice, Firenze 2003. Principi<br />

Ecologici per la Progettazione del Paesaggio, Franco Angeli, Milano 2007.<br />

Finotto F., Monaco R., Servente G., Un modello per la valutazione della produzione e della<br />

<strong>di</strong>ffusività <strong>di</strong> energia biologica in un sistema ambientale, to be printed on Scienze Regionali (Ital. J.<br />

of Regional Sci.), 2010.<br />

Forman R. T. T. 1995. Land Mosaics. The ecology of landscape and regions. Cambridge:<br />

Cambridge Press.<br />

Gobattoni F., Lauro G., Leone A., Monaco R., Pelorosso R., 2010. A simulation method for the<br />

stability analysis of landscape scenarios by using a NetLogo application in GIS environment. EGU<br />

Conference, 3-7 May, Vienna, Austria.<br />

Hurlbert A. H. and Jetz W. 2010. More than “More In<strong>di</strong>viduals”: The Non-equivalence of Area and<br />

Energy in the Scaling of Species Richness. Am Nat 2010. Vol. 176, pp. 000–000 DOI:<br />

10.1086/650723.<br />

Ingegnoli V. 2002. Landscape Ecology: A Widening Foundation. New York-Berlin: Springer-<br />

Verlag.<br />

Ingegnoli V., Forman R.F., 2002. Landscape Ecology: A Widening Foundation, Springer-Werlag,<br />

New York.<br />

Lauro G., Lisi M., Monaco R., 2008. Bifurcation analysis of a dynamical model for an ecological<br />

system. La <strong>Matematica</strong> e le sue Applicazioni, n. 15. on-line ISSN 1974-305X.<br />

Naveh Z., Liebermann A. 1984. Landscape ecology: theory and application. Springer-Werlag, New<br />

York.<br />

Pelorosso R., Leone A., Boccia L. 2009. Land cover and land use change in the Italian central<br />

Apennines: A comparison of assessment methods. Applied Geography 29:35–48.<br />

Perry L.W., 2002. Landscape, space and equilibrium: shifting viewpoints. Progress in Physical<br />

Geography, 26 (3):339-359.<br />

Petit C.et al., 2008. Landscape Analysis and Visualisation-Spatial Models for Natural Resources<br />

and Planning. Springer,<br />

Tscharntke T., Klein A. M., Kruess A., Steffan-Dewenter I.and Thies C. 2005. Landscape<br />

perspectives on agricultural intensification and bio<strong>di</strong>versity – ecosystem service management.<br />

Ecology Letters, 8 (8):857-874.<br />

Turner M. G., Romme W. H., Gardnerl R. H., O’Neill R. V. and Kratz T. K. 1993. A revised<br />

concept of landscape equilibrium: Disturbance and stability on scaled landscapes. Landscape<br />

Ecology, 8(3):213-227.<br />

Turner M.G., Gardner R.H., 1990. Quantitative methods in Landscape Ecology. Springer-Werlag,<br />

New York.<br />

Vermaat, J.E., Eppink, F., Van den Bergh, J.C.J.M., Barendregt, A. & Van Belle, J. 2005. Matching<br />

of scales in spatial economic and ecological analysis. Ecol. Econ., 52, 229-237.<br />

Willemen L., Verburg P.H., Hein L., Van Mensvoort M. E. F. (2008). Spatial characterization of<br />

landscape functions. Landscape and Urban Planning, 88:34-43.

Hooray! Your file is uploaded and ready to be published.

Saved successfully!

Ooh no, something went wrong!