MILIOLIDAE - Acta Palaeontologica Polonica

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W11b Fig. 23. Ontogeny and dimorphism of QuinquelocuHna triangularis d'Orbigny, Lower Tortonian, Wieliczka; 1-MS, micro generation; 2 - QS, micro generation, 3-6 - QS, mega I generation, 7-11- QS, of triloculine appearance, mega II generation; a front view, b apertural view, c cross-section.
MILIOLIDAE FROM MIOCENE OF POLAND 65 1968. Quitiqueloculina triangularis d'Orbigny; J. P. Margerel, p. 16, PI. 5, Figs 21-22 (non PI. 6, Fig. 1). 1969b. Quinqueloculina akneriana akneriana d'Orbigny; A. K. Bogdanowich, p. 86, PI. 2, Fig. 8a, b. Material. - Over 1000 QS and 20 MS (CoIl. No. F-115, Sec. No.139 142,315,317-319,346,347,349-351). Dimensions. QS - L 0.8-1.25; B 0.5-1.05; T 0.5-0.8; MS - L 1.65-3.5; B. 0.8-1.25; T 0.4-0.8. Description. - QS broadly oval, thick, high-triangular in cross-section, periphery subacute; chambers broad, flat on one side and slightly inflated on the other; middle chamber large, broad, prominent; juvenile stage may be triloculine; MS large, flat, with broadly rounded periphery and indistinct middle chambers; sutures slightly depressed, indistinct; wall thin in QS, thick in MS; surface smooth, polished; aperture oval, not large, with a simple, bifid tooth. Variability. The shape of massiline forms varies from circular to broadly oval, whereas the quinqueloculine and triloculine forms are always the same shape. A distinctive feature of the massiline forms is their rounded periphery, while the quinqueloculine forms show a particular deformation of chambers. which are increasingly more inflated on th9 front side than on the back side. This is well seen in cross-sections (Textfig. 23). Dimorphism. Massiline forms belong generally to the micro generation (Text-fig. 23/1), quinqueloculine forms to the micro and mega I generations (Text-fig. 23/2-6) and triloculine forms to the mega II generation (Text-fig. 2317-11). The morphological differentiation of generations is not marked except for the size of the proloculus and the number of chambers. PD: micro - 20-30 ~l, mega I - 100-110 fl, mega II 200-300 fl. Remarks. - Massiline forms show the same shape and internal structure as Miliolina podolica Didkovsky, 1959 does. It is clearly seen from the cross-section of this species given by its author (op. cit., Text-fig. 2) and from those of our specimens, that the juvenile stage has its internal structure identical with that of Quinqueloculina triangularis d'Orbigny. The only difference betwen them is the thicker wall in the juvenile stage of the massiline forms of Q. triangularis which indicates that there is a possibility for calcium carbonate to accumulate on the chamber walls of the juvenile stage while successive massiline chambers are being built. The quinqueloculine stage removed from the inside of the massiline form corresponds morphologically to Q. triangularis (Textfig. 24). It should be mentioned that both the species have been described from strata of the same age, namely Q. triangularis from the lithothamnian facies of the Upper Tortonian of Nussdorf in the Vienna Basin, and M. podolica from the clay facies of the Upper Tortonian in Podolia. 5 Acta Palaeontologica nr 1/74
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