MILIOLIDAE - Acta Palaeontologica Polonica

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56 EWA LUCZKOWSKA Distribution. - Poland: Lower Tortonian (Grabki Duze, Karsy, Korytnica, L~ki Dolne, Wieliczka). Quinqueloculina parakneriana sp.n. (PI. VII, Figs la, b, 2a, b, 3a, b, 4a, b; Text-fig. 19) Holotypus: PI. VII, Fig. 4a, b. Paratypi: PI. VII, Figs la, b, 2a, b, 3a, b. Locus typicus: Korytnica near J~drzej6w. Stratum typicum: Lower Tortonian (Badenian), gray clay marls. Derivatio nominis: parakneriana - close to Q. a..kneriana. Diagnosis. - Test oval with broadly rounded periphery, thick and massive; walls inflated, tending to become flat. Material. - 30 QS and 17 MS, (ColI. No. F-I09, Sec. No. 232, 273, 352, 353, 356, 357). Dimensions: Holotype MS length 2.2 mm, breadth 1.8 mm, thickness 0.75 mm; Paratypes QS length 1.1-1.7 mm, breadth 0.8-1.2 mm, thickness 0.6-0.75 mm; MS length 1.65-1.75 mm, breadth 1.25-1.45 mm, thickness 0.65-0.75 mm. Description. - Quinqueloculine stage narrowly oval, tapering at both ends, periphery subrounded; chambers of uniform width, arcuate, sides nearly flat, middle chamber broad, much elongated, with prominent rounded edge, fifth chamber very small, occasionally invisible; massiline stage oval, large, robust, thick, slightly flattened, periphery nearly rounded with faintly marked edge; chambers broad, inflated, middle chamber elongate, narrowly oval, previous chambers slightly visible; sutures distinct, depressed; wall thick; surface smooth, polished; aperture large, elongate oval, with a flexuose margin surrounded with a thickened rim and furnished with a long bifid tooth. Variability. Typical characters of the species are the oval shape of test, subrounded periphery and depressed sutures. The convexity of chambers is variable in massiline and large quinqueloculine forms, in which there is a tendency for the walls to become flattened and, as a result, a slightly pronounced keel is formed on one or two last chambers in some specimens. Dimorphism. Only small quinqueloculine forms belong to the megalospheric II generation, having a proloculus 150 !! in diameter (Text-fig. 19/5). The other massiline and quinqueloculine forms studied are of the microspheric generation, the proloculus of which is 10-20 II in diameter (Text-fig. 19/1-4). Ontogeny. - As can be seen in sections, the roundeness of the periphery appears rather in adult specimens, being mostly subrounded in the juvenile stage. Most specimens of this species are quinqueloculine and they generally attain a large size. The massiline stage is rare. Remarks. - Massiline forms differ from Quinqueloculina akneriana d'Orbigny MS in having a thicker and more massive test, a more slender
<strong>MILIOLIDAE</strong> FROM MIOCENE OF POLAND 57 shape, more inflated walls, and a subrounded periphery with a slightly marked keel; quinqueloculine forms are more slender, having a lowtriangular shape in cross-section and no tendency to the deformation of the chamber walls. The two species are otherwise undobtedly related. They ~lb®®~~ rJQ~q~~~,~, Fig. 19. Ontogeny and dimorphism of Quinqueloculina parakneriana n. sp., Lower Tortonian, Korytnica; 1,2 - MS of quinqueloculine appearance, micro generation; 3,4-QS, micro generation; 5 - QS, mega I generation; a front view, b apertural view, c cross-section. differ from Q. pseudobuchiana sp.n. in their more slender shape and rounded or subrounded, not acute or keeled, periphery. The massiline forms most resemble Q. araucana d'Orbigny from Recent seas, differ however in being twice as large and in having a more oval shape and a broader and bigger middle chamber. The quinqueloculine forms differ from Q. laevigata d'Orbigny in broader chambers and in more flattened walls. Distribution. - Poland: Tortonian (Benczyn, Gliwice Stare, Grabki Duze, Iwkowa, Karsy, Korytnica, L~ki Dolne). Czechoslovakia: Lower Tortonian, Zidlochovice. Quinqueloculina peregrina d'Orbigny, 1846 (PI. -II, Figs la-c, 2a, b; Text-fig. 15/1-3) 1846. Quinqueloculina peregrina d'Orbigny; A. d'Orbigny, p. 292, PI. 19, Figs 1-3. Material. - 20 QS and 130 MS (ColI. No. F-llO, Sec. No. 103-107). Dimensions: QS - L 1.10-1.25; B 0.7-0.85; T 0.35-0.5; MS - L 1.40-1.85; B 1.05-1.55; T 0.4-0.5.
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