MILIOLIDAE - Acta Palaeontologica Polonica

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36 EWA LUCZKOWSKA lmm Fig. 3. Morphological variations and ontogeny of Quinqueloculina akneriana d'Orbigny, Lower Tortonian, Wieliczka; 1,2 - MS; 3 - transitional form; 4,5 - QS; a front view, b apertural view. lmm @@.@. (~~m@':J® ~b~ 4b 5b 6b ii~~~@b~ 2c 3c 4c 5c 6c lb Fig. 4. Ontogeny and dimorphism of Quinqueloculina akneriana d'Orbigny, Lower Tortonian, Wieliczka; 1 - MS, micro generation, a front view, b cross-section; 2 - MS, mega I generation; 3,5,6 - QS, mega I generation; 4 a front view, b apertural view, c cross-section. feature is the deformation of chambers in most of the quinqueloculine specimens, in which the frontal wall of each succeeding chamber is more convex than that of the back wall of the chamber. Dimorphism. 3) Both QS and MS belong to the micro and mega I ge- 3) The abbrevations: P - proloculus, PD - proloculus diameter.
<strong>MILIOLIDAE</strong> FROM MIOCENE OF POLAND 37 nerations (Text-fig. 4). The mega II generation has not been found. PD: micro 20!l' mega I 70-160fl. Ontogeny. - Only a few MS specimens have been found beside a large number of QS ones. The massiline stage was probably rarely attained in this species and mature individuals remained quinqueloculine. If we expose the juvenile stage by removing the external chambers of a specimen of "Quinqueloculina pauperata d'Orbigny" appearance, we receive a form of Quinqueloculina akneriana d'Orbigny appearance. (Text-fig. 5). lmm @a ~b Fig. 5. Quinqueloculina akneriana d'Orbigny and its juvenile stage, Lower Tortonian, Wieliczka; 1 - MS; 2 - QS from inside MS; a front view, b apertural view. The morphology of tests derived from the same sample indicates the presence of transitional forms between Q. akneriana and "Q. pauperata" (Text-fig. 3). Remarks. - Our specimens are identical with the topotypes from Baden, but differ somewhat from d'Orbigny's illustration from 1846 in having a less rounded periphery. There is great morphological resemblance and probably also a relationship in internal structure between Q. akneriana and Q. triangularis d'Orbigny. Q. akneriana QS differs from Q. triangularis QS in its more· slender shape and more distinct and depressed sutures. The aperture shape and the internal structure are identical in these species. The difference in the shape of tooth noted by d'Orbigny is insignificant as both species may have either a simple or a bifid tooth. The MS of both species is however indistinguishable, even in slightly damaged specimens. The QS of the mega II generation of both species is probably indistinguishable too, hence they are hard to identify. In spite of this conformity the quinqueloculine stage of these two species can be easily distinguished from each other on the basis of the test shape and therefore they are not regarded here as synonyms, as they are by some authors (e.g. Marks 1951, Verhoeve 1970). In the materials from Baden (Sooss) and Wieliczka both species are common and they differ evidently one from another. Both authors mentioned above synonymize a number of other species of d'Orbigny, Reuss, Bornemann and even Q. seminulum
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