MILIOLIDAE - Acta Palaeontologica Polonica

168 EWA LUCZKOWSKA
'iHeM JlHllIb Y HeKOTOpbIX 3K3eMnJlRpOB ,l\aHHOrO BH,l\a). Ba:lKHbIM npH3HaKOM RBJlReTCR
TaK:lKe pa3HaR cPOpMa YCTbR H 3y6a, OC06eHHO Y TaKHX POAOB KaK Cycloforina, Miliolinella,
Pyrgo, Pyrgoella, Varidentella. O'ieHb peAKO BCTpe'iaIOTCR BHAbI, Y
KOTOpbIX
MOPcPOJlOrH'ieCKHe npH3HaKH H3MeHRIOTCR B
He3Ha'iHTeJlbHOH CTeneHH.
OHTorEHE3
Ha6J1IO,l\eHHR Ce'ieHHH pRAa cPOPM pa3HbIx BH,l\OB B nOpR,l\Ke OT MJlaAeH'ieCKHX
CTaAHH no B3pOCJlbIe HJlH CTap'ieCKHe, HJlH :lKe H3y'leHHe CTa,l\HH pa3BHTHR OTAeJlb­
HbIX 3K3eMnJlRpOB n03BOJlReT CAeJlaTb CJleAyro~He 3aKJlIO'ieHHR:
1) BeJlH'iHHa paKOBHHbI He 3aBHCHT o'r KOJlH'ieCTBa KaMep H 06YCJlOBJleHa, npe:lK,l\e
Bcero, BeJlH'iHHOH Ha'iaJlbHOH KaMepbI. 'tJeM 60JlbWe Ha'laJlbHaR KaMepa, TeM MeHbwe
KaMep y
oc06eH O,l\HHaKOBOH BeJlH'iHHbI. CJleAOBaTeJlbHO, MJlaAeH'ieCKaR CTaAHR reHepa~HH
Mera I HJlH Mera II MO:lKeT HMeTb TaKyIO :lKe BeJlH'iHHY, KaK H
B3pOCJlaR CTa,l\HR
reHepa~HH MHKpO. 2) B
COCTaBe po,l\a Quinqueloculina, 06pa3YIO~ero B3POCJlYIO (HJlH
CTap'iecKyro?) MaccHJlHHOBYIO CTaAHIO, Ha6J1IOAaIOTCR TaKHe BH,l\bI, y KOTOpbIX eAHH­
CTBeHHO KBHHKBeJlOKYJlHHOBaR CTaAHR RBJlReTCR pewaIO~HM npH3HaKoM npHHaAJle:lK­
HOCTH K
AaHHOMY BHAY, TaK KaK cP0PMbI MaCCHJlHHOBbIX CTa,l\HH MorYT 6bITb BeCbMa
CXOAHbI Y pa3HbIx BHAoB. 3TO MO:lKHO paccMaTpHBaTb ABORKO: JlH60 cy~ecTByeT POA
Massilina, a pa3HbIe cPOPMbI MJlaAeH'ieCKOH cTaAHH, Ha6J1IOAaIO~HeCR He'. Ce'ieHHRX
CXOAHbIX cPOPM, CBHAeTeJlbCTBYIOT 0 HX nOJlHcPHJleTH'ieCKOM npOHCXO:lKAeHHH, JlH60 :lKe
POA Massilina npeACTaBJlReT JlHWb oHToreHeTH'lecKYIO CTa,l\HIO pOAa Quinqueloculina,
H30MOPcPHYIO Y pa3HbIx BH,l\OB. B pa60Te npHHRTa BTopaR TO'iKa 3peHHR (CM. 'iaCTb I).
3) Ha Ce'ieHHRX 3K3eMnJlRpOB pRAa pOAOB MO:lKHO Ha6J1IO,l\aTb, 'iTO MJlaAeH'leCKafl
cTaAHfl HeKOTopbIX BH,l\OB HMeeT CXO,l\HbIH 06JH1K, a
HHAHBHAyaJlbHbIe npH3HaKH npoflBJlflIOTCfl
TOJlbKO JlHWb nocJle CcPOpMMpOBaHHfl HeCKOJlbKHX Ha~IaJlbHbIX KaMep (HanpHMep,
PO,l\bI Cycloforina, LachlanelZa, Triloculina). OTCIO,l\a CJleAyeT, 'ITO 3a HCKJlIO­
'ieHHeM pOAa Quinqueloculina MJlaAeH'leCKaR cTaAHfl APyrHx POAOB JlHWeHa MH,l\HBM­
,l\yaJlbHbIX AMarHOCTM'leCKMX npM3HaKoB. 4) BHYTPM p0,l\a Sinuloculina npMcYTcTByIOT
BMAbI, MJla,l\eH'leCKafl cTaAHfl KOTOpbIX HaCTOJlbKO OTJlH'laeTCfl OT B3POCJlOH CTa,l\MH,
'iTO MX npMHMMaIOT B
Ka'leCTBe pa3HhIX BHAOB.
'tJTO KacaeTCfl cPOPMhI yCThfl M 3y6a, TO y HeKOTopbIX pO,l\OB, HanpMMep y VaridentelZa,
Hauerina, Miliola OHM HMeIOT pa3HbIf1 BMA B MJlaAeH'leCKOH M B3POCJlOH
CTaAMflX.
PO,l(CTBEHHbIE OPMbJ
Ha6J1IOAeHHR IIOKa3aJlH, 'ITO cpeAM MHJlMOJlM,l\ MMo~eHa IIoJlhwH MO:lKHO 3aMeTHTh
TaKHe rpynnhI cPOPM BHyTpH oAHoro pO,l\a, p0,l\CTBeHHaR CBR3h KOTOPbIX BbIpa:lKeHa
TaK CXOAHbIM cTpoeHHeM MJla,l\eH'ieCKOW CTa,l\HH, KaK M HeKoTopbIMM MOPcPOJlOrM'ie­
CKHMH npM3HaKaMM B3POCJlOW CTa,l\MM, a TaK:lKe CXO,l\HhIM TMIIOM CTeHKW.
TIPOME:lKYTO'l:HbIE OPMbI ME:lK,l(Y PO,l(AMH
TpY,l\HOCTh onpeAeJleHHfl HeKoTophIX POAOB MHJlMOJlMA 06YCJlOBJleHa, B
OCHOBHOM,
TeM, 'ITO cy~eCTBYIOT BHAhI CO
"CMewaHHhIMM" AMarHOCTH'ieCKHMM IIpM3HaKaMM, BbI-
3bIBaIO~WMM HeyBepeHHOCTh B OTHeceHHH HX K OIIpeAeJleHHOMY POAY. K TaKMM npH-