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K - College of Natural Resources - University of California, Berkeley

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Descriptions <strong>of</strong> all outbreaks and issues specific to each dataset are outlined later in the<br />

SOM.<br />

For the Poisson, geometric and negative binomial models, the maximum<br />

likelihood estimate <strong>of</strong> the basic reproductive number (R0 or R) was the sample mean<br />

(Rice 1995). For the negative binomial distribution, the shape parameter k was also<br />

estimated by maximum likelihood estimation (mle). A numerical algorithm was used<br />

for maximization, which required that an upper bound be set for the range <strong>of</strong> k sampled.<br />

This upper bound was set to k=1000, and when this bound was returned as the mle<br />

estimate then ˆ was set to infinity, because a NegB(R0,k=1000) distribution is<br />

k mle<br />

indistinguishable in practice from Poisson(R0).<br />

Having computed the maximum likelihood scores for each dataset, we compared<br />

the Poisson, geometric and negative binomial models using Akaike’s information<br />

criterion (AIC) (Anderson et al. 2000):<br />

where ( L(<br />

ˆ | data )<br />

( L(<br />

ˆ | data ) K<br />

AIC = -2 ln θ + 2<br />

ln θ is the log-likelihood maximized over the unknown parameters (θ),<br />

given the model and the data, and K is the number <strong>of</strong> parameters estimated in the model.<br />

Because some <strong>of</strong> our datasets are small, we used the modified criterion AICc, which<br />

reduces to the conventional expression as sample size N becomes larger (Anderson et al.<br />

2000):<br />

AIC c<br />

= -2 ln θ<br />

( ( ˆ<br />

2K<br />

) ( K + 1)<br />

L | data + 2K<br />

+<br />

129<br />

N − K −1

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