Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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Distribution: Rio de 1 aneiro, Brazil (Fig. 14). Habitat: unknown. Common name: unknown. Economic use: unknown. Beilschmiedia stricta is known to me only by the type collection. This species appears to be similar to B. curviramea and B. fluminensis. About the difference between B. stricta and these two species, see the notes under these two species respectively. 26. Beilschmiedia taubertiana (Schwacke & Mez) Kostennans, Rec. Trav. Bot. Neerl. 35: 863. 1938. Hufelandia taubertiana Schwacke et Mez, Arb. Bot. Gart. Breslau 1: 108. 1892. TYPE: Brazil. Minas Geraes: Rio Novo, 1890 (fl., collection in B with fl. & fr.), Alaujo 7047 (holotype, B seen on photograph; isotypes, P, RB). Tree, height unknown. Terminal buds pubescent with erect long straight hairs. Twigs terete, younger ones compressed, pubescent with erect long straight or slightly wavy hairs, less densely when old. Leaves opposite; petioles 0.8-1 em long, canaliculate above, pubescent with erect long hairs, slightly discolored with twigs; blades firmly chartaceous, elliptic, 10-14 x 4-5 em; base obtuse to round, not inrolled, apex acute; upper leaf surface glabrous, lower leaf surface sparsely pubescent with erect, long straight to slightly wavy hairs; midrib immersed above, raised below, secondary veins 9-14 pairs, slightly raised or almost imn1ersed above, raised below, tertiary veins not percurrent, ramification coarse, square-like with free veinlets, slightly raised on both sides. Inflorescences axillary, racemose, ca. 1.5 em long, pubescent with long erect hairs, with ca. 5 flowers per inflorescence; flower pedicels of the lateral divisions 0.5-1 mm long, pedicels of the central flowers up to 4 mn1long. Flowers ca. 3 mm long, ca. 3 rnn1 in diam.; tepals 6, equal, ovate, 1.9-2 mm long, 1.3-1.5 mm wide, pubescent with erect long wavy hairs outside, sparsely pubescent with appressed to erect hairs inside; stamens 9, outer 6 filaments ca. 0.3 mm long, innermost 3 filaments ca. 0.4 mm long, filaments pubescent, outer six anthers 1.2-1.3 mm long, innermost anthers 0. 9-1.1 mm long, all the anthers 2-celled, apex of the anthers obtuse to truncate, anthers pubescent, glands of the innermost 3 stamens globose; staminodia 3, cordate in outline, ca. 0.6 mm long; pistil ca. 1.2 n1illlong, glabrous, ovary shorter than and gradually narrowed into the style; receptacle densely pubescent with erect 82
hairs. Fruits ellipsoid, ca. 5 x 3 em, the surface warty; infructescence axis ca. 3 mm in diam., fruit pedicels thickened to ca. 5 mn1 in diam. below the fruit, fruit pedicels not constricted at the base, covered with lenticels and concolorous with the infructescence axis. Phenology: unknown. Distribution: Minas Gerais, Brazil (Fig. 14). Habitat: unknown. Common name: unknown. Economic use: unknown. Additional specimen examined: Brazil. Minas Gerais: Rio Novo, 1889 (sterile), Arm~jo 5 (RB). Beilschmiedia taubertiana has opposite leaves and coarse ramification as all the Brazilian Beilschmiedia species do. But this species is easily distinguished fron1 the other Brazilian species by the erect, long straight pubescence on the terminal buds and twigs, and by the lower leaf surfaces sparsely pubescent with same kind of indument. 27. B eilscluniedia tovarensis (Meissner) Sa. Nishida comb. nov. Hufelandia tovarensis Meissner, D. C. Prodr. 15: 65. 1864. SYNTYPES: Colombia. locality unknown, Karsten 88 (B seen on photograph). Venezuela. Aragua: Tovar, ca. 1920 malt., Aug. 28, 1854 (fl.), Fendler 1094 (NY). Aniba pseudo-coto (Rushy) Kostermans, Rec. Trav. Bot. Neerl. 35: 872. 1938. Ocotea pseudo-colo Rusby, Bull. Torrey Bot. Club 49: 261. 1922. TYPE: Bolivia. La Paz: mountains S of Huachi, near the Cochabamba River, 750-900 malt., Sep. 9, 1921 (young fl .), White 1051 (holotype, NY seen on photograph; isotypes, AFS not seen, C, G-DEL not seen, GH not seen, K, US seen on photograph). Tree, to 40 m tall. Terminal buds pubescent with appressed straight hairs. Twigs terete, slightly angular when young, pubescent with appressed straight hairs, less densely when older. Leaves alternate; petioles (0.5) 1-4.2 em long, flat to canaliculate above, glabrous or pubescent with appressed straight hairs, concolorous with twigs; blades coriaceous to fim1ly chartaceous, ovate to broad elliptic, rarely broad oblanceolate, (5) 8-20 83
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KURENAI : Kyoto University Researc
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Contents Abstract Chapter 1 Chapter
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Chapter 1 General Introduction Laur
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Chapter 2 Revisional study of neotr
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Akkermans", if it is not a type, Be
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Straightness of the indument son1et
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Staminodia in the fourth whorl are
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elongs not to B. ova/is but to B. a
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Many species occur fron1 premontane
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Although I did not make an intensiv
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12a. Inflorescences racen1ose, more
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1. Beilscluniedia alloiophylla (Rus
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slopeS of creek, NE slope of Altos
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Rica. This group appears to have re
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anay, and he stated that the Colotn
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Phenology: collected in flower in M
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angustifolia has pubescent pistils,
Page 35 and 36: The specimen with a label saying "B
Page 37 and 38: 23, 1990 (fr.), Estinoza45 (CR, INB
Page 39 and 40: Beilschmiedia rohliana Lasser, Bol.
Page 41 and 42: (fl.), Haber 11837 (MO); La Peii.a
Page 43 and 44: (F); Con. Ceylan, path La Esmeralda
Page 45 and 46: Colombia (Cuatrecasas 22220, 22438)
Page 47 and 48: locality, Nov. 29- Dec. 18, 1964 (s
Page 49 and 50: Among Brazilian Beifschmiedia speci
Page 51 and 52: van der Werff et al. 12951 (holotyp
Page 53 and 54: inside; starnens 9, filaments ca. 0
Page 55 and 56: ecause the flowers of the type coll
Page 57 and 58: This species also shares a similar
Page 59 and 60: synonym of B . sulcata, supposedly
Page 61 and 62: Additional specimen examined: Brazi
Page 63 and 64: Cristobal leaving main Hwy 100.8 mi
Page 65 and 66: pubescent with erect, curly to wavy
Page 67 and 68: Tree, to 20 m tall. Terminal buds p
Page 69 and 70: 16 x (5) 6-10 em; base obtuse to cu
Page 71 and 72: Alajuela: Canton de Alfaro Ruiz, Pa
Page 73 and 74: There are several collections from
Page 75 and 76: Spiritus (Las Villas): same mountai
Page 77 and 78: (fl.), Ekman Hl4565 (MO), Apr. 3, 1
Page 79 and 80: 316, 1906, Little Jr. 6293, Aguilar
Page 81 and 82: Cerro Hueco, ca. 700 malt., Feb. 4,
Page 83 and 84: ecause there is a distribution gap
Page 85: Beilschmiedia immersinervis also ap
Page 89 and 90: Additional specimens examined: Cost
Page 91 and 92: 2, 1994 (young fl.), van der Werff
Page 93 and 94: In the illustration of Ruiz and Pav
Page 95 and 96: Table 1. List of the Beilschmiedia
Page 97 and 98: Legends of figures Fig. 1. Five typ
Page 99: 4 A B c . w ; . -)1 . ..... J.f D f
Page 102 and 103: o Beilschmiedia berteroana * B. bre
Page 104 and 105: o Beilschmiedia hexanrhera * B. hon
Page 106 and 107: 0 o Beilschmiedia ovaLioides *B. ov
Page 108 and 109: C::::J.o•' :: ~ ·a ' \) ' ,o 0 .
Page 110 and 111: Chapter 3. Leaf Anatomy of neotropi
Page 112 and 113: 3. Leaf sections The technique used
Page 114 and 115: 16), B. ovalis (Fig. 18), B. peruiu
Page 116 and 117: smooth. However, in B. angustielfip
Page 118 and 119: uneven-sized subsidiary cells. This
Page 120 and 121: Midrib. The most distinct character
Page 122 and 123: This grouping is more or less suppo
Page 124 and 125: In leaf sectional study, neotropica
Page 126 and 127: Species of group 2 have leaves oppo
Page 128 and 129: Table 1. List of samples for venati
Page 130 and 131: 0\ Table 2. Continued. B. emarginat
Page 132 and 133: Legends of figures Fig. 1-25. Venat
Page 134 and 135: Fig. 122-143. Midrib sections of ne
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2 132
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10 1 1 12 \ 14 134
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References Allen, C. K. 1945. Studi
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Meissner, C. F. 1864. Lauraceae. Pp
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