Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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ordering hills, Jun. 24, 1957 (sterile), Stern173 (NY); Can a region, on trail above Cana leading to ridgeofPirre Massif, T45'N 7T45'W, 800-900 m alt., May 5, 1990 (immature & mature fr.), McPherson 15040 (F, MO). Panama: Sendero de Interpretacion, 1km E of camp in the reserveforest of INRENARE, 8°40'N 79°55'W, 800-900 m alt., Mar. 24, 1994 (fl.), Correa 10408 (MO); same locality, 800-900 m alt., Mar. 30, 1994 (fl.), Correa 11039 (MO); same locality, 800-900 m alt., Jun. 29, 1995 (immature fr.), Correa 11164 (MO). Venezuela. Merida: Highway Paname1icana, between rivers Gavilan & R. Perdido, 150-500 malt., Jun. 2, 1960 (fr.), VeilLon 11112 (MO). Miranda: Districto Paez, road between San Juan- Montevideo, 10°04'-06'N, 65"45'-47'W, 400 malt., Sep. 7, 1977 (young fi.), Gonz&ez 1363 (MO). Colombia. Antioquia: Municipio San Luis, stream "La Cristalina", 6°N 74°45'W, 730-770 m alt., May. 22, 1987 (immature fr.), Guillermo 933 (MO). Narino: Espriella, Fumaeo, Jun. 22, 1951 (fr.), Castaneda 2820 (F). Ecuador. Esmeraldas: N of San Lorenzo, Panadero, Silv. Pare. Panadero, 40 m alt., Apr. 30, 1966 (immature fr. or sterile), lanse 279 (NY, US (sterile)); Rio Onzole, 90 malt., Jul. 14, 1966 (immature fr.), Jdtiva316 (NY), (immature fr.), JG!iva1 096 (NY, US); San Lorenzo, near rail road, right of way 3 km SE, ca. 9 malt., Apr. 20, 1943 (young fl. & young fr.), Little, Jr. 6293 (F, NY), US); same locality, 4? m alt., Apr. 20, 1943 (young fi. & young fr.), Little, Jr. 6294 (MO, NY, US). Napo: 5 k~ N of Coca, El Chuncho Floristic Reserve, 00°25'S 7TOl'W, 250m alt., May23, 1993 (young fl.), Palacios 10794 (MO); Canton Archidona, S slopes ofYolcan Sumaco, Hollfn-Loreto road,40 km, Huamani, 00o43'S 7T36'W, 1200 malt., Oct. 19, 1989 (fl.), Neil/9066 (MO), (fl.), Palacios4655 (MO); Estaci6n Biol6gica Jatun Sacha, Rfo Napo, 8 km E of Misahuallf, 01 °04'S 77°36'W, 200m alt., Jun. 23-27, 1987 (young fl.), Cer6n 1662 (MO); Orellana, Yasunf Nacional Park, road & pipeline of Maxus in construchtion, 53-60 km, 0°48'S 76°28'W, 230m alt., Oct. 6, 1993 (fr.), Dik 626 (MO). Beilschtniedia pendula is the only Beilschmiedia species distributed in the West Indies. There is actually a collection reported fron1 the West Indies (Jan1aica) belonging to another Beilschmiedia species (B . curviramea). But there is no more collection of this species from the West Indies and I suspect that the locality of the collection was n1islabeled. Beilschmiedia pendula has alternate leaves and fme ramification. Among the species of the san1e phyllotaxis and ran1ification, B. pendula is one of the 1nost inconspicuous species, often confused with B. mexicana, B. costaricensis, and B. tovarensis. These four species share similar pubescence type, leaf shape (except for a group of B. tovarensis which has ovate-broad elliptic leaves), flower structure and fruit shape. Fron1 B. mexicann and B. costaricensis, B. pendula can be distinguished best by its ramification almost in1n1ersed on upper leaf surfaces and glaucous lower leaf surfaces (usually, B. tnexicann and B. costaricensis have conspicuously raised ramification on both leaf surfaces and non-glaucous leaves). However, there are some exceptional collections of B. pendula fron1 Costa Rica and Ecuador which have non glaucous leaves (lanse 279, Jmiva 74
316, 1906, Little Jr. 6293, Aguilar 2619, Yasuda 1315 and 1317), and some exceptional collections of B. mexicana which have the ran1ification almost immersed above or glaucous leaves (about the exceptional collections of B. mexicana, see the notes under B. tnexicana). From B. costaricensis, B. pendula also differs by its fruit pedicels constricted at the base. And leaf apex of B. pendula is usually acuminate and different from the acute to obtuse leaf apex of B. mexicana and B. costaricensis, but there are some exceptional collections in B. pendula with obtuse leaf apex (see below). About the difference between B. pendula and B. tovarensis, see the notes under B. tovarensis. Leaves of B. pendula are usually relatively small and narrow elliptic-oblanceolate, but there are some exceptional collections with large and wide leaves, especially from Puerto Rico and Martinique. Leaf apex of these collections are also different from the typical collections, the typical collections usually have distinctively acuminate leaf apex, but the collections from Puerto Rico and Martinique tend to have obtuse leaf apex. 22. Beilschrniedia rigida (Mez) Kosterrnans, Rec. Trav. Bot. Neerl. 35: 856. 1938. Hufelandia rigida Mez in Taubert, Bot. Jahrb. Syst. 17: 519. 1893. SYNTYPES: Brazil. Rio de Janeiro: Alto Macahe de Nova Friburgo, Oct.-Nov. (fl.), Glaziou 19790 (B not seen, BM, C, F, LE not seen, P); Glaziou 19793 (B not seen, BR, G-BOIS not seen, K, LE not seen, NY, P, US not seen). Tree, height unknown. Tem1inal buds pubescent with appressed short straight hairs. Twigs terete, sparsely pubescent with appressed short straight hairs or almost glabrous. Leaves opposite; petioles 1.5-3.5 em long, flat above, slightly discolored with twigs, glabrous; blades coriaceous, obovate, 14-22 x 7-11 em; base decurrent, flat to slightly inrolled to below, apex round to obtuse; leaf surface glabrous on the both sides, not known whether glaucous or not; midrib in11nersed above, raised below, secondary veins 9-12 pairs, slightly raised above, raised below, tertiary veins not percurrent, ramification coarse, square-like with free veinlets, slightly raised on both sides. Inflorescences axillary or clustered around the terminal buds, paniculate, 4-12 (18) em long, sparsely pubescent with appressed to erect hairs or almost glabrous, with 10-30 (55) flowers per inflorescence; flower pedicels of the lateral divisions 1.5-2 mm long, pedicels of the central flowers up to 5 mm long. Flowers 3-3.5 mm long, ca. 3.5 111m in diam.; tepals 6, equal, ovate, 1.5-2.3 mm 75
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KURENAI : Kyoto University Researc
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Contents Abstract Chapter 1 Chapter
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Chapter 1 General Introduction Laur
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Chapter 2 Revisional study of neotr
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Akkermans", if it is not a type, Be
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Straightness of the indument son1et
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Staminodia in the fourth whorl are
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elongs not to B. ova/is but to B. a
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Many species occur fron1 premontane
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Although I did not make an intensiv
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12a. Inflorescences racen1ose, more
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1. Beilscluniedia alloiophylla (Rus
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slopeS of creek, NE slope of Altos
Page 27 and 28: Rica. This group appears to have re
Page 29 and 30: anay, and he stated that the Colotn
Page 31 and 32: Phenology: collected in flower in M
Page 33 and 34: angustifolia has pubescent pistils,
Page 35 and 36: The specimen with a label saying "B
Page 37 and 38: 23, 1990 (fr.), Estinoza45 (CR, INB
Page 39 and 40: Beilschmiedia rohliana Lasser, Bol.
Page 41 and 42: (fl.), Haber 11837 (MO); La Peii.a
Page 43 and 44: (F); Con. Ceylan, path La Esmeralda
Page 45 and 46: Colombia (Cuatrecasas 22220, 22438)
Page 47 and 48: locality, Nov. 29- Dec. 18, 1964 (s
Page 49 and 50: Among Brazilian Beifschmiedia speci
Page 51 and 52: van der Werff et al. 12951 (holotyp
Page 53 and 54: inside; starnens 9, filaments ca. 0
Page 55 and 56: ecause the flowers of the type coll
Page 57 and 58: This species also shares a similar
Page 59 and 60: synonym of B . sulcata, supposedly
Page 61 and 62: Additional specimen examined: Brazi
Page 63 and 64: Cristobal leaving main Hwy 100.8 mi
Page 65 and 66: pubescent with erect, curly to wavy
Page 67 and 68: Tree, to 20 m tall. Terminal buds p
Page 69 and 70: 16 x (5) 6-10 em; base obtuse to cu
Page 71 and 72: Alajuela: Canton de Alfaro Ruiz, Pa
Page 73 and 74: There are several collections from
Page 75 and 76: Spiritus (Las Villas): same mountai
Page 77: (fl.), Ekman Hl4565 (MO), Apr. 3, 1
Page 81 and 82: Cerro Hueco, ca. 700 malt., Feb. 4,
Page 83 and 84: ecause there is a distribution gap
Page 85 and 86: Beilschmiedia immersinervis also ap
Page 87 and 88: hairs. Fruits ellipsoid, ca. 5 x 3
Page 89 and 90: Additional specimens examined: Cost
Page 91 and 92: 2, 1994 (young fl.), van der Werff
Page 93 and 94: In the illustration of Ruiz and Pav
Page 95 and 96: Table 1. List of the Beilschmiedia
Page 97 and 98: Legends of figures Fig. 1. Five typ
Page 99: 4 A B c . w ; . -)1 . ..... J.f D f
Page 102 and 103: o Beilschmiedia berteroana * B. bre
Page 104 and 105: o Beilschmiedia hexanrhera * B. hon
Page 106 and 107: 0 o Beilschmiedia ovaLioides *B. ov
Page 108 and 109: C::::J.o•' :: ~ ·a ' \) ' ,o 0 .
Page 110 and 111: Chapter 3. Leaf Anatomy of neotropi
Page 112 and 113: 3. Leaf sections The technique used
Page 114 and 115: 16), B. ovalis (Fig. 18), B. peruiu
Page 116 and 117: smooth. However, in B. angustielfip
Page 118 and 119: uneven-sized subsidiary cells. This
Page 120 and 121: Midrib. The most distinct character
Page 122 and 123: This grouping is more or less suppo
Page 124 and 125: In leaf sectional study, neotropica
Page 126 and 127: Species of group 2 have leaves oppo
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Table 1. List of samples for venati
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0\ Table 2. Continued. B. emarginat
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Legends of figures Fig. 1-25. Venat
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Fig. 122-143. Midrib sections of ne
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2 132
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10 1 1 12 \ 14 134
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136
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138
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140
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References Allen, C. K. 1945. Studi
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Meissner, C. F. 1864. Lauraceae. Pp
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