inflorescence; flower pedicels <strong>of</strong> the lateral divisions 0.5-1 (1.5) nun long, pedicels <strong>of</strong> the central flowers up to 4 mn1long. Flowers greenish yellow to whitish, 2-3 mm long, 2.3-2.8 mm in diam.; tepals 6, equal, ovate, 1.2-1.7 mm long, 0.8-1.4 mm wide, sparsely pubescent with appressed to erect hairs on both sides; stamens 9, outer six filaments 0.3-0.4 mm long, innermost three filaments 0.4-0.6 mm long, filaments pubescent, anthers ca. 0. 8 mm, all the anthers 2-celled, apex <strong>of</strong> the anthers obtuse-acute to truncate, glabrous, glands <strong>of</strong> the innermost stamens globose; staminodia 3, cordate in outline, ca. 0.8 mm long; pistil ca. 1.5 mm long, glabrous or rarely sparsely pubescent, ovary almost as long as and gradually narrower into the style; receptacle pubescent with appressed hairs. Fruits ellipsoid, black, shiny, ca. 2-3.5 (4.5) x ca. 1.5 em, the surface smooth; infructescence axis ca. 2 mm in diam., fruit pedicels constricted at the base, slightly thickened below the fruit, infructescences not covered with lenticels and the fruit pedicel concolorous with the infructescence axis. Phenology: collected in flower from February to November, in mature fruit from March to November. Distribution: Honduras, Nicaragua, Costa Rica, Pana1na, West Indies, Venezuela, Colombia and Ecuador, 0-1680m alt. (Fig. 13). Habitat: montane broad-leafed forest, tropical wet forest, sometimes on limestone. Comn1on name: "Mulato" and "Came de doucella" in Cuba, "Lauries formis" in St. Lucia, "Vacarita" in Panama, "Carabali" and "Cobalongo mach" in Venezuela. Economic use: reported to be useful as a tin1ber in Dominica, Martinique, Montserrat and St. Lucia. Additional specimens examined: Cuba. Camagliey: Lorna del Gato and Vicinity Cobre Range <strong>of</strong> Sierra Maestra, Jul. 11-Aug. 14, 1921 (immature fr.), Leon 10451 (NY). Cienfuegos (Las Villas): SE <strong>of</strong> Cumanayagua, Sierra de San Juan, 3-400m alt., Jul. 1941, (fl.), Howard 5695 (F, MO, NY, U). Guatanmo (Oriente): E. Cuba, Monte Verde, 1856-7 (fl. & immature fr.), Wright 485 (BM, BR, MO, NY); same locality, Sep. 1859-Jan. 1860 (immature fr.), Wright 1402 (K, MO, NY), (fl.), Wright 1403 (1402 & 1402, rarely 1414, on same sheets) (K, MO), (immature fr.), Wright 1414 (BR, MO, NY); Rfo Purial, 200m alt., May, 1889 (immature fr.), Eggers 5434 (P); locality unknown, 1856-7 (fr.), Wright 486 (BR, NY, MO (sterile)). Holguin (Oriente): Pierra deNipe, at Rfo Piedras, 500 malt., Aug. 24, 1915 (immaturefr.),Ekman6402 (F, NY, US); Sierra de Nigre (Negro?), at Rio Pilato, ca. 350m alt., Mar. 14, 1915 (sterile), Ekman 5009 (NY). Sancti Spiritus (El Porvenir): Santa Clara, Mts. Trinidad, 650-750 malt., Mar. 9, 1910 (fl.), Britton5320 (NY). Sancti 70
Spiritus (Las Villas): same mountains, San Bias-Buenos Aires, Gaviiias, Sep. 18, 1941 (young fr.), Gonzales 173 (B, BM, NY). Sancti Spiritus (Santa Clara): Mts. Banao, May., 1920 (immature fr.), Luma?650 (NY); same locality, 950 malt., Jul. 30, 1918 (immature fr.), Leon 7975 (NY); Sierra deGavilanes, Mts. Sancti-Spiritus, Aug. 1-2, 1916 (immature fr.), Leon 6612 (NY). Santiago de Cuba (Oriente): S side <strong>of</strong> the crest <strong>of</strong> the Sierra Maestra, La Bayamesa, W <strong>of</strong> Aserradero San Antonio de los Cumbres, 1400-1500 malt., Jan. 23-24, 1956 (young fl.), Morton 9563 (US); Sierra Maestra S <strong>of</strong> Yara in headwaters <strong>of</strong> Rio Yara, Alto de Naranjo, Trail to Pico Turquino, 76°52'W, 900-950m alt., Jul. 9, 1985 (fl. & young fr.), Gentry50952 (MO, NY); S. Oriente and Pico Turquino, Sauks <strong>of</strong> arroyo del Cristo, Aug. 10, 1922 (immature fr.), Ekmanl4826 (NY). Villa Clara (Las Villas): St. Domingo, 600 m alt., May. 24, 1887 (fl.), Eggers 2087 (NY). Villa Clara (Santa Clara): Buenos Aires, Trinidad Hills, 750-1150 malt., Mar. 8, 1929 (immature fr.), Jack 7016 (NY); Buenos Aires, Las Lagunas, ca. 750 rn alt., Dec. 5, 1928 (immature fr.), Jack6806 (F (2 sheets), NY, US). Jamaica. Clarendon, Along road between Ritchies & Balcarres, 900 malt., Nov. 10, 1973 (immature fr.), Proctor33607 (MO); same locality, 900 m alt., Apr. 24, 1974 (fl.), Proctor33791 (NY); same locality, 900 malt., Oct. 5, 1977 (immature fr.), Proctor 37316 (MO); same locality, 900 malt., Jun. 12, 1976 (old fl. & young fr.), Thorne48212 (MO. NY); Manchester, Sep. 21-23, 1908 (immature fr.), Britton 3691 (NY); Manderille, ca. 630 m alt., Sept., 21, 1908 (immature fr.), Harris] 0594 (F, NY, US); near Siver Hill, Set. 25, 1894 (sterile),Harris5286 (BM); Scher Hill Galp., ca. 1050m alt., Aug. 10, 1895 (fl.), Harris 5843 (BM, F, NY, US); Spanish River, Aug. 24, 1894 (young fr.), Harris5326 (BM); Troy, Sep. 13-18,1906 (fl.), Britton660 (F, NY); Troy, ca. 480 malt., Sep. 18, 1906 (fr.), Harris 9447 (NY); Whitfield Hall, Feb. 1, 1895 (immaturefr.),Harris5575 (BM,F). Haiti. Ouest: Gros Cheval, Momes des Commissaires, 1400 m alt., Mar. 17, 1942 (immature fr.), Holdridge I 053 (BM, F, MO, NY (2 sheets), US). Ouest or Artibonte: Corail, Camp #1, ca. 600 malt., Jul. 16-17, 1905 (inunature fr.), Nash1035 (NY). Sud: Grand'Anse-Sud limit, Massif de La Hotte, 13.6 km N <strong>of</strong> Camp Perrin on road between Roseaux andJeremie, ''TombeauCheval", 18°23'N73°53'W, 720m alt., Nov. 15,1982 (irrunaturefr.),Zanoni 24313 (MO); Massif deJa Hotte, Pic Macaya National Park, on hills S <strong>of</strong> MorneFormon, near Ville Formon, 1200-1250 m alt, Feb. 1, 1984 (immature fr.), Judd3983 (NY); Massif de la Hotte, N Tranchant, nearGodet, ca. 1600 malt., Sep. 19, 1924 (fl.), EkmanH1950 (K, US). Locality unknown: Jun. 19, 1964 (fl.), Augusto 1569 (NY). Dominica. Barahona: ''El Gajo" ca. 7 km from the carreteradeCabral a Polo, through the way to "Entrada de Cortico" ("Monteada Nueva"), 18°07.5'N 71 °13 .5'W, 1400 malt., Jan. 18, 1986 (immature fr.), Zanoni 36009 (MO, NY); "Entrada de Cortico" ("Monteada Nueva"), above the batey de cafe "Monte Nueva", on the peak <strong>of</strong> Ia Lorna Trocha de Pey, 18°07.5'N 71 °13.5'W, 1400 malt., Jan. 14, 1988 (immature fr.), Zanoni40676 (MO, NY, U); Mt. Nueva, Cafia Brava, 1300 malt., Apr. 24, 1976 (young fl.), Liogier 25140 (NY); same locality (S . <strong>of</strong> Cabral), 1300 malt., Jun. 15, 1968 (fl.), Liogier 11658 (NY); Mt. Nueva, 6.3 km from Cruce de El Puerto-Monteado-Nueva, 3.4 km E <strong>of</strong>Cortico, 18° 10'N 71 °15'W, 1390 m alt., Apr. 11, 1985 (fr.), Gentry 50705 (MO); Sierra de Baoruco, Lomas Ia Trocha dePey, above Monteada Nueva, 1400 m alt., Oct. 16, 1982 (immature fr.), Proctor 391 OJ (MO); Sierra de Baoruco; 4 km above a small village "EntradadeCortico", enroute to El Gajo, 18°07.5N 71 o13 .5'W, 1320-1340 malt., Mar. 25, 1990 (fr.), Zanoni 44246 (MO, NY); Sierra de Baoruco,onthehill toMomeLaJo, ca.8km N<strong>of</strong>Poloporcarnino, 18°08'N7lol8'W, 1200-1300malt.,Feb. 71
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KURENAI : Kyoto University Researc
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Contents Abstract Chapter 1 Chapter
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Chapter 1 General Introduction Laur
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Chapter 2 Revisional study of neotr
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Akkermans", if it is not a type, Be
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Straightness of the indument son1et
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Staminodia in the fourth whorl are
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elongs not to B. ova/is but to B. a
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Many species occur fron1 premontane
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Although I did not make an intensiv
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12a. Inflorescences racen1ose, more
- Page 23 and 24: 1. Beilscluniedia alloiophylla (Rus
- Page 25 and 26: slopeS of creek, NE slope of Altos
- Page 27 and 28: Rica. This group appears to have re
- Page 29 and 30: anay, and he stated that the Colotn
- Page 31 and 32: Phenology: collected in flower in M
- Page 33 and 34: angustifolia has pubescent pistils,
- Page 35 and 36: The specimen with a label saying "B
- Page 37 and 38: 23, 1990 (fr.), Estinoza45 (CR, INB
- Page 39 and 40: Beilschmiedia rohliana Lasser, Bol.
- Page 41 and 42: (fl.), Haber 11837 (MO); La Peii.a
- Page 43 and 44: (F); Con. Ceylan, path La Esmeralda
- Page 45 and 46: Colombia (Cuatrecasas 22220, 22438)
- Page 47 and 48: locality, Nov. 29- Dec. 18, 1964 (s
- Page 49 and 50: Among Brazilian Beifschmiedia speci
- Page 51 and 52: van der Werff et al. 12951 (holotyp
- Page 53 and 54: inside; starnens 9, filaments ca. 0
- Page 55 and 56: ecause the flowers of the type coll
- Page 57 and 58: This species also shares a similar
- Page 59 and 60: synonym of B . sulcata, supposedly
- Page 61 and 62: Additional specimen examined: Brazi
- Page 63 and 64: Cristobal leaving main Hwy 100.8 mi
- Page 65 and 66: pubescent with erect, curly to wavy
- Page 67 and 68: Tree, to 20 m tall. Terminal buds p
- Page 69 and 70: 16 x (5) 6-10 em; base obtuse to cu
- Page 71 and 72: Alajuela: Canton de Alfaro Ruiz, Pa
- Page 73: There are several collections from
- Page 77 and 78: (fl.), Ekman Hl4565 (MO), Apr. 3, 1
- Page 79 and 80: 316, 1906, Little Jr. 6293, Aguilar
- Page 81 and 82: Cerro Hueco, ca. 700 malt., Feb. 4,
- Page 83 and 84: ecause there is a distribution gap
- Page 85 and 86: Beilschmiedia immersinervis also ap
- Page 87 and 88: hairs. Fruits ellipsoid, ca. 5 x 3
- Page 89 and 90: Additional specimens examined: Cost
- Page 91 and 92: 2, 1994 (young fl.), van der Werff
- Page 93 and 94: In the illustration of Ruiz and Pav
- Page 95 and 96: Table 1. List of the Beilschmiedia
- Page 97 and 98: Legends of figures Fig. 1. Five typ
- Page 99: 4 A B c . w ; . -)1 . ..... J.f D f
- Page 102 and 103: o Beilschmiedia berteroana * B. bre
- Page 104 and 105: o Beilschmiedia hexanrhera * B. hon
- Page 106 and 107: 0 o Beilschmiedia ovaLioides *B. ov
- Page 108 and 109: C::::J.o•' :: ~ ·a ' \) ' ,o 0 .
- Page 110 and 111: Chapter 3. Leaf Anatomy of neotropi
- Page 112 and 113: 3. Leaf sections The technique used
- Page 114 and 115: 16), B. ovalis (Fig. 18), B. peruiu
- Page 116 and 117: smooth. However, in B. angustielfip
- Page 118 and 119: uneven-sized subsidiary cells. This
- Page 120 and 121: Midrib. The most distinct character
- Page 122 and 123: This grouping is more or less suppo
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In leaf sectional study, neotropica
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Species of group 2 have leaves oppo
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Table 1. List of samples for venati
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0\ Table 2. Continued. B. emarginat
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Legends of figures Fig. 1-25. Venat
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Fig. 122-143. Midrib sections of ne
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2 132
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10 1 1 12 \ 14 134
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136
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138
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140
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142
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144
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146
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148
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150
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152
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154
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References Allen, C. K. 1945. Studi
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Meissner, C. F. 1864. Lauraceae. Pp