Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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SE ofTotontepec, 250m alt.? (mistake for2500 m ?), Aug. 9, 1977 (fl.), Sousa 7871 (UC); Districto Villa Alta, 8.1 km N of Maravillas, road between Talea de Castro and Yalina, 2370 malt., May 15, 1983 (fl. & a part of fr.), Torres 2947 (F, MEXU, MO); Municipio San Felipe Usila, 7.3 km S (179°) of Santa Cruz Tepetotutla, 17"40'23"N 96o33'28"W, 2220 malt., May. 15, 1994 (fr.), Rincon 419 (MEXU); same municipio, 7.6 km S (179°) of Santa Cruz Tepetotutla, 1T40'13"N 96°33'28'W, 2240 malt., May. 16, 1994 (fl.), Rincon 433 (MEXU). Beilschmiedia ovalioides appears to be close to B. ovalis. 1l1e two species share very similar pubescence type, ramification, flower structure and fruit shape. But B. ovaLioides has longer petioles (more than 2.0 em long), lower leaf surfaces pubescent with curly hairs, secondary veins more than 11 pairs and conspicuously percurrent tertiary veins, while B. ovalis has shorter petioles (less than 1.5 cn1 long), glabrous lower leaf surfaces, secondary veins less than 9 pairs and tertiary veins not conspicuously percurrent. Those characters are not strong individually, but the cOinbination of then1 separate these two species clearly. Additionally, B. ovalioides tends to have larger (n1ore than 11 em long) and broadly elliptic to obovate leaves, whereas B. ovalis usually has smaller (less than 10 em) and ovate or elliptic leaves. Both of the species occur above 1800 m in elevation, but the distribution has some disjunction; B. ovalioides has been collected only in Mexico, whereas B. oval is has been collected in Honduras, Costa Rica and Pana1na. One specimen of B. ovalioides is reported from 250 n1 alt., but considering the locality, this altitude is likely a mistake for 2500 malt. There is a collection fron1 Chiapas which has pubescent lower leaf surfaces, but has shorter petioles (1 .5-2.0 em long) and less secondary veins (8-10) than the typical B. ovalioides (Martfnez 19456). Because the leaves have abnormal apices, I place this collection in B. ovalioides, assuming that the leaves are son1ewhat damaged. 20. Beilschmiedia oYalis (Blake) C. K. Allen, J. Arnold Arbor. 26:418. 1945. Hufelandia ovalis Blake in Jour. Wash. Acad. Sci. 9 :461. 1919. TYPE: Costa Rica. Alajuela: Volcan de Poas, 2300 n1 alt., (fl.), Mar. 31, 1907, Pittier2040 (holotype, US; isotype, F). Beilschmiedia austin-smithii (Standley) C. K. Allen, J. Arnold Arbor. 25: 418. 1945. Persea austin-smithii Standley, Field. Mus. Publ. Bot. 18:1552. 1938. Type: Costa Rica. 66
Alajuela: Canton de Alfaro Ruiz, Palmira, (fl.), Apr. 30, 1937, Smith 4168 (holotype: F). Tree, to 30m tall. Terminal buds pubescent with erect short curly hairs. Twigs terete to angular, densely to sparsely pubescent with short curly hairs, less denser to glabrous when old. Leaves alternate; petioles 1-1.5 cn1 long, flat above, glabrous or rarely pubescent with short curly hairs, concolorous with twigs; blades coriaceous, ovate to broadly elliptic, (4) 5- 10 (14) x (3) 4-8 (10.5) em; base obtuse to round, not inrolled, apex obtuse; leaf surface glabrous on both sides, rarely partly pubescent with erect curly hairs when young, lower surface glaucous; midrib in1mersed above, raised below; secondary veins 5-8 (9) pairs, immersed above, raised below; tertiary veins not percurrent or weakly percurrent, ramification fine, areolate without free veinlets, immersed or slightly raised above, slightly to conspicuously raised below. Inflorescences axillary or clustered on the terminal buds, paniculate, (3) 5-15 cn1long, pubescent with erect hairs, with 20-50 flowers per inflorescence; flower pedicels of the lateral divisions 0.5-1.2 mm long, pedicels of the central flowers up to 2 n1m long. Flowers yellowish, 2.5-3.2 mm long, 2.9-3.7 mm in diam.; tepals 6, equal, ovate to elliptic, (1.4) 1.8-2.3 mm long, 1-1.7 nun wide; pubescent with erect hairs on both sides; stamens 9, outer six filaments 0.3-0.5 mm long, innermost three filaments 0.5-0.7 mm long, filaments pubescent, anthers ca. 1 mm long, all of them 2- celled, apex of the anthers more or less obtuse, apex of the anthers glabrous, glands of the innermost three stan1ens globose; stan1inodia cordate in outline, 0.5-1 rnm long; pistil 1.3- 1.7 mm long, glabrous, ovary almost as long as or longer than and gradually narrowed into the style; receptacle relatively densely pubescent with long, erect to almost appressed hairs. Fruits (known only in immature stage) roundish, green when immature, ca. 3.5 x ca. 3.5 em (or bigger), the surface smooth; fruit pedicels ca. 5 mm in dian1., thickened to ca. 10 mm in diam. below the fruit; pedicels not constricted at the base, concolorous with the lower part. Phenology: collected in flower from February to May, and in September and Noven1ber, in mature fruit in November. Distribution: Honduras, Costa Rica, Panama, in 1800-2800 malt. (Fig. 12). Habitat: cloud forest. Common nan1e: "Aguacate negro'' in Costa Rica, "Aguacatillo" in Panama. 67
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KURENAI : Kyoto University Researc
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Contents Abstract Chapter 1 Chapter
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Chapter 1 General Introduction Laur
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Chapter 2 Revisional study of neotr
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Akkermans", if it is not a type, Be
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Straightness of the indument son1et
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Staminodia in the fourth whorl are
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elongs not to B. ova/is but to B. a
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Many species occur fron1 premontane
Page 19 and 20: Although I did not make an intensiv
Page 21 and 22: 12a. Inflorescences racen1ose, more
Page 23 and 24: 1. Beilscluniedia alloiophylla (Rus
Page 25 and 26: slopeS of creek, NE slope of Altos
Page 27 and 28: Rica. This group appears to have re
Page 29 and 30: anay, and he stated that the Colotn
Page 31 and 32: Phenology: collected in flower in M
Page 33 and 34: angustifolia has pubescent pistils,
Page 35 and 36: The specimen with a label saying "B
Page 37 and 38: 23, 1990 (fr.), Estinoza45 (CR, INB
Page 39 and 40: Beilschmiedia rohliana Lasser, Bol.
Page 41 and 42: (fl.), Haber 11837 (MO); La Peii.a
Page 43 and 44: (F); Con. Ceylan, path La Esmeralda
Page 45 and 46: Colombia (Cuatrecasas 22220, 22438)
Page 47 and 48: locality, Nov. 29- Dec. 18, 1964 (s
Page 49 and 50: Among Brazilian Beifschmiedia speci
Page 51 and 52: van der Werff et al. 12951 (holotyp
Page 53 and 54: inside; starnens 9, filaments ca. 0
Page 55 and 56: ecause the flowers of the type coll
Page 57 and 58: This species also shares a similar
Page 59 and 60: synonym of B . sulcata, supposedly
Page 61 and 62: Additional specimen examined: Brazi
Page 63 and 64: Cristobal leaving main Hwy 100.8 mi
Page 65 and 66: pubescent with erect, curly to wavy
Page 67 and 68: Tree, to 20 m tall. Terminal buds p
Page 69: 16 x (5) 6-10 em; base obtuse to cu
Page 73 and 74: There are several collections from
Page 75 and 76: Spiritus (Las Villas): same mountai
Page 77 and 78: (fl.), Ekman Hl4565 (MO), Apr. 3, 1
Page 79 and 80: 316, 1906, Little Jr. 6293, Aguilar
Page 81 and 82: Cerro Hueco, ca. 700 malt., Feb. 4,
Page 83 and 84: ecause there is a distribution gap
Page 85 and 86: Beilschmiedia immersinervis also ap
Page 87 and 88: hairs. Fruits ellipsoid, ca. 5 x 3
Page 89 and 90: Additional specimens examined: Cost
Page 91 and 92: 2, 1994 (young fl.), van der Werff
Page 93 and 94: In the illustration of Ruiz and Pav
Page 95 and 96: Table 1. List of the Beilschmiedia
Page 97 and 98: Legends of figures Fig. 1. Five typ
Page 99: 4 A B c . w ; . -)1 . ..... J.f D f
Page 102 and 103: o Beilschmiedia berteroana * B. bre
Page 104 and 105: o Beilschmiedia hexanrhera * B. hon
Page 106 and 107: 0 o Beilschmiedia ovaLioides *B. ov
Page 108 and 109: C::::J.o•' :: ~ ·a ' \) ' ,o 0 .
Page 110 and 111: Chapter 3. Leaf Anatomy of neotropi
Page 112 and 113: 3. Leaf sections The technique used
Page 114 and 115: 16), B. ovalis (Fig. 18), B. peruiu
Page 116 and 117: smooth. However, in B. angustielfip
Page 118 and 119: uneven-sized subsidiary cells. This
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Midrib. The most distinct character
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This grouping is more or less suppo
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In leaf sectional study, neotropica
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Species of group 2 have leaves oppo
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Table 1. List of samples for venati
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0\ Table 2. Continued. B. emarginat
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Legends of figures Fig. 1-25. Venat
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Fig. 122-143. Midrib sections of ne
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2 132
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10 1 1 12 \ 14 134
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136
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138
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140
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References Allen, C. K. 1945. Studi
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Meissner, C. F. 1864. Lauraceae. Pp
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