Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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Haber 11092 (INB, MO); P. I. La Amistad; Cordillera de Talamanca; Estaci6n Biol6gica Las Altura, 8°57' 15''N 83°50' lO'W, 1580 m alt., Mar.7 .1996 (sterile), Yasuda 1307 (INB). Burger and van der Werff (1990) reported that a few collections which he placed in Beilschmiedia ovalis had obovate leaves, atypical for B. ovalis, and he commented that they might belong to another, undescribed, species. I have also observed several collections which share similar pubescence type and leaf texture with B. oval is but have obovate leaves. They have been collected froxn lower elevation than where B. oval is occurs, and they consistently have ellipsoid fruits, quite different fron1 the spherical fruits of B. oval is. Although I have not seen a significant difference in flower between these collections and B. ova/is (I have seen only one flowering collection with obovate leaves, which has less pubescent inflorescences and slightly sn1aller flowers than B. ovalis), I think obovate leaves and ellipsoid fruits are strong enough characters to distinguish the collections from B. oval is. In addition to the two characters above, this new species has the tertiary veins conspicuously percurrent, which is rarely seen in B. oval is. Usually, B. obovatifolia has the ran1ification conspicuously raised below and the fruit pedicels slightly thickened below the fruits. There are two collections from Costa Rica (Burger 12184 and Dayton 3126) which have obovate leaves, but I keep the collections separate from this species as indeterminate, because the collections have almost immersed ramification and the fruits of the former collection have narrow pedicels. 19. Beilschmiedia ovalioides Sa. Nishida sp. nov. TYPE: Mexico. Oaxaca: Distrito de Ixtlan, Sierra de Juarez, enroute from Xiacui to Talea de Castro, ca. 25 kn1 SSW of Talea fron1 the route, 2750 malt., Apr. 19, 1982 (fl. & fr.), Lorence 4078 (holotype, MEXU; isotype, F, MO). A Beilschmiedia ovalis petiolis longioribus et foliis subtus puberulis differt. Tree, to 35 m tall. Terminal buds densely pubescent with rusty, erect short curly hairs. Twigs terete, slightly angular when young, densely pubescent with rusty, erect short curly hairs. Leaves alternate; petioles ( 1. 7) 2-3 em long, flat above, rarely slightly canaliculate above, pubescent with rusty short curly hairs, less densely when old, concolorous with twigs; blades coriaceous, broadly elliptic to obovate, sometimes slightly asymmetric, (8) 11- 64
16 x (5) 6-10 em; base obtuse to cuneate, not inrolled, apex acute to acuminate; upper leaf surface glabrous, lower leaf surface puberulent with short curly hairs, lower surface usually glaucous; midrib and secondary veins immersed above, raised below, secondary veins (8) 11-14 pairs, tertiary veins percurrent, ramification fine, areolate without free veinlets, immersed above, slightly raised or immersed below (ramification pattern visible below). Inflorescences in the axils of the leaves near the terminal buds, paniculate, ( 4.5) 6-12 em long, pubescent with rusty, erect short curly hairs, with 40-80 flowers per inflorescence; flower pedicels of the lateral divisions 0.5-0.7 mm long, pedicels of the central flowers up to 1 mm long. Flowers greenish-yellow, ca. 2.8 mm long, ca. 3 mm in diam.; tepals 6, equal, ovate, 1.8-2.2 mm long, 1.2-1.5 (2) nlffi wide, densely pubescent with erect hairs outside, sparsely pubescent with erect curly hairs to almost glabrous inside; stamens 9, filaments 0.3- 0.5 mm long, pubescent, anthers ca. 0.9 mm long, all the anthers 2-celled, apex of the anthers obtuse to truncate, glabrous, glands of innermost three stamens globose; staminodia 3, cordate in outline, ca. 0.7 mm; pistil ca. 1.5 mm long, glabrous, ovary as long as the and gradually narrowed into the style; receptacle pubescent with long, appressed to erect hairs. Fruits roundish, black, 3-3.5 x 3-4 em, the surface smooth; infructescence axis ca. 5 mm in diam., fruit pedicels thickened to 10 nm1 in diam below the fruit; fruit pedicels not constricted at the base, covered with few lenticels and slightly darker than the infructescence axis. Phenology: collected in flower in May and August, in mature fruit in February, April and November. Distribution: Mexico, 1850-2750 n1 alt. (Fig. 12). Habitat: montane mesophyll forest. Common name: unknown. Econon1ic use: unknown. Additional specimens exan1ined: Mexico. Chiapas: Municipio Union Juarez, in El Yolcan Tacana by a road from Talquian to the top of the volcano, along the border with Guatemala, 2200-2700 malt., Feb. 4, 1987 (fr.), Martinezl9456 (MO). Oaxaca: Sierra de Juarez, ca. 25km SSW ofTaleadeCastro, or3km N of the junction to Yalina, limit of the district of Ixtlan & Villa Alta, 2750 malt., May 31, 1983 (fl.), Cedilfo2372 (F, MEXU, MO); W of Municipio San Felipe Usila, 8 km straightS from Santa Cruz Tepetotutla, 1 T40'06''N 96°33'24'W, 2395 malt., Mar. 31, 1994 (young fl.), Gallardo 1007 (MEXU); Districto Choapam, roadfromTotontepecto Comaltepec, 5 km NE of Totontepec, 1850 m alt., Nov. 11, 1983 (fr.), Lorence4376 (F, MEXU, MO); 20 km 65
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KURENAI : Kyoto University Researc
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Contents Abstract Chapter 1 Chapter
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Chapter 1 General Introduction Laur
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Chapter 2 Revisional study of neotr
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Akkermans", if it is not a type, Be
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Straightness of the indument son1et
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Staminodia in the fourth whorl are
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elongs not to B. ova/is but to B. a
Page 17 and 18: Many species occur fron1 premontane
Page 19 and 20: Although I did not make an intensiv
Page 21 and 22: 12a. Inflorescences racen1ose, more
Page 23 and 24: 1. Beilscluniedia alloiophylla (Rus
Page 25 and 26: slopeS of creek, NE slope of Altos
Page 27 and 28: Rica. This group appears to have re
Page 29 and 30: anay, and he stated that the Colotn
Page 31 and 32: Phenology: collected in flower in M
Page 33 and 34: angustifolia has pubescent pistils,
Page 35 and 36: The specimen with a label saying "B
Page 37 and 38: 23, 1990 (fr.), Estinoza45 (CR, INB
Page 39 and 40: Beilschmiedia rohliana Lasser, Bol.
Page 41 and 42: (fl.), Haber 11837 (MO); La Peii.a
Page 43 and 44: (F); Con. Ceylan, path La Esmeralda
Page 45 and 46: Colombia (Cuatrecasas 22220, 22438)
Page 47 and 48: locality, Nov. 29- Dec. 18, 1964 (s
Page 49 and 50: Among Brazilian Beifschmiedia speci
Page 51 and 52: van der Werff et al. 12951 (holotyp
Page 53 and 54: inside; starnens 9, filaments ca. 0
Page 55 and 56: ecause the flowers of the type coll
Page 57 and 58: This species also shares a similar
Page 59 and 60: synonym of B . sulcata, supposedly
Page 61 and 62: Additional specimen examined: Brazi
Page 63 and 64: Cristobal leaving main Hwy 100.8 mi
Page 65 and 66: pubescent with erect, curly to wavy
Page 67: Tree, to 20 m tall. Terminal buds p
Page 71 and 72: Alajuela: Canton de Alfaro Ruiz, Pa
Page 73 and 74: There are several collections from
Page 75 and 76: Spiritus (Las Villas): same mountai
Page 77 and 78: (fl.), Ekman Hl4565 (MO), Apr. 3, 1
Page 79 and 80: 316, 1906, Little Jr. 6293, Aguilar
Page 81 and 82: Cerro Hueco, ca. 700 malt., Feb. 4,
Page 83 and 84: ecause there is a distribution gap
Page 85 and 86: Beilschmiedia immersinervis also ap
Page 87 and 88: hairs. Fruits ellipsoid, ca. 5 x 3
Page 89 and 90: Additional specimens examined: Cost
Page 91 and 92: 2, 1994 (young fl.), van der Werff
Page 93 and 94: In the illustration of Ruiz and Pav
Page 95 and 96: Table 1. List of the Beilschmiedia
Page 97 and 98: Legends of figures Fig. 1. Five typ
Page 99: 4 A B c . w ; . -)1 . ..... J.f D f
Page 102 and 103: o Beilschmiedia berteroana * B. bre
Page 104 and 105: o Beilschmiedia hexanrhera * B. hon
Page 106 and 107: 0 o Beilschmiedia ovaLioides *B. ov
Page 108 and 109: C::::J.o•' :: ~ ·a ' \) ' ,o 0 .
Page 110 and 111: Chapter 3. Leaf Anatomy of neotropi
Page 112 and 113: 3. Leaf sections The technique used
Page 114 and 115: 16), B. ovalis (Fig. 18), B. peruiu
Page 116 and 117: smooth. However, in B. angustielfip
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uneven-sized subsidiary cells. This
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Midrib. The most distinct character
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This grouping is more or less suppo
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In leaf sectional study, neotropica
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Species of group 2 have leaves oppo
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Table 1. List of samples for venati
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0\ Table 2. Continued. B. emarginat
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Legends of figures Fig. 1-25. Venat
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Fig. 122-143. Midrib sections of ne
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2 132
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10 1 1 12 \ 14 134
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136
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138
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140
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References Allen, C. K. 1945. Studi
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Meissner, C. F. 1864. Lauraceae. Pp
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