Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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15. Beilsch1niedia linharesensis Sa. Nishida & van der Werff. TYPE: Brazil. Espirito Santo: Linhares, Reserva Florestal da Porto Seguro- CVRD/BA, Aceiro, 2100 km, left side, Oct. 26, 1988 (fl.), Farias 243 (holotype, MO). Beilschmiedia rigida simi lis, a qua floribus minoribus et depresse globosis differt. Tree, to ca. 30 m tall. Terminal buds pubescent with appressed straight hairs. Twigs terete, compressed when young, sparsely to densely pubescent with appressed straight hairs. Leaves opposite; petioles 1.5-2 cn1 long, flat above, slightly darker than twigs, glabrous; blades coriaceous, obovate to elliptic, (5.5) 8-14 x (3) 4-5 em; base cuneate, inrolled to below (entire margin slightly inrolled to below), apex round to obtuse; leaf surface glabrous on the both sides, not glaucous; midrib immersed or slightly impressed above, raised below, secondary veins 8-12 (14) pairs, slightly raised on both sides, tertiary veins not percurrent, ramification coarse, square-like with free veinlets, almost immersed on both sides. Inflorescences axillary, paniculate with few branching, 2.5-3.5 em long, relatively densely pubescent with erect straight hairs, with ca. 15 flowers per inflorescence; flower pedicels of the lateral division ca. 0.5 mm long, pedicels of the central flowers up to 1 mm long. Flowers depressed-globose, ca. 1.6 (2) mm long, ca. 2.2 mm in diam.; tepals 6, equal, curved toward inside, broad ovate, ca. 0.5 mm long, ca. 1 mm wide, sparsely pubescent with erect hairs outside, sparsely pubescent with erect hairs to almost glabrous inside, stamens 9, outer six filaments ca. 0.1 mm long, innermost three filaments ca. 0.2 mm long, filaments pubescent, anthers ca. 0.5 mm long, all anthers 2-celled, apex of the anthers obtuse to truncate, apex of the anthers pubescent, glands on innermost three stamens globose; staminodia 3, cordate in outline, ca. 0.3 mm long; pistil ca. 1.2 mm long, glabrous, ovary as long as and gradually narrowed into the style; receptacle pubescent with appressed to erect hairs. Fruits unknown. Phenology: collected in flower in October. Distribution: Espirito Santo, Brazil (Fig. 11). Habitat: unknown. Common name: unknown. Economic use: unknown. 56
Additional specimen examined: Brazil. Espirito Santo: Reserva Florestal de CVRD. Linhares, Est. CINCO-FOLHAS, 1320 km right side, Sep. 16, 1982 (young fl.), Folli 409 (MO). Although I have only one collection in n1ature flower and two collections in flower buds, the small depressed-globose flowers of this species are unique among neotropical Beilschmiedia species and distinguished from the other species clearly. Beilschmiedia linharesensis shares sinlilar phyllotaxis, ramification pattern, pubescence type and leaf shape with B. rig ida. 16. Beilschmiedia ntexicana (Mez) Kostermans, in Rec. Trav. Bot. Neerl. 35: 846. 1938. Hufelandia mexicana Mez, Jahrb. Konig!. Bot. Gart. Berl. 5: 20. 1889. SYNTYPES: Mexico. Veracruz: Dos Puentes, Feb. (fl.), Liebman 712 (= ''Lauraceae N. 20") (C not seen, Knot seen, US not seen); Mirador, Feb., 1842 (fr.), Liebman 711 (= ''Lauraceae N. 16" ?) (US), same locality, Mar., 1842 (immature fr.), Liebman 711 (="LauraceaeN. 19" ?) (C); Liebman 711 (="LauraceaeN . 16"or ''LauraceaeN. 19")(P not seen); Liebman 714 (= "Lauraceae N. 21 ") (C not seen, LE not seen); Totutla, Jan., 1843 (fr.), Liebman 713 (= "Lauraceae N . 17, 18") (C, US). Orizaba, 1853 (fr.), F. Muller 1460 (B not seen, K-2 sheets, LE not seen, W not seen). Linociera areolata Lundell, Amer. Midi. Naturalist 23: 176. 1940. TYPE: Mexico. Hidalgo: above Chapulhuacan, 1300 malt., Jul. 12, 1937 (fr.), Lundell & Lundell7165 (holotype, AFS not seen; isotype, F) . Tree, to 25 m tall. Terminal buds pubescent with appressed hairs, rarely almost glabrous. Twigs terete, compressed when young, pubescent with appressed hairs or glabrescent, less densely to glabrous when older. Leaves alternate, petioles 0.5-1.5 (2) em long, flat to canaliculate above, glabrous, rarely pubescent with appressed hairs, concolorous with twigs; blades chartaceous, elliptic, sometimes asymmetric, 5-13 (19) x 3-8 (10) em; base cuneate, not inrolled, apex acute to obtuse, rarely acuminate; leaf surface glabrous on both sides, lower leaf surface not glaucous, or very rarely glaucous; midrib and secondary veins almost immersed or slightly raised above, raised below, secondary veins in 6-10 (II) pairs, ramification raised on both size, very rarely immersed above(? see below), 57
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KURENAI : Kyoto University Researc
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Contents Abstract Chapter 1 Chapter
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Chapter 1 General Introduction Laur
Page 7 and 8:
Chapter 2 Revisional study of neotr
Page 9 and 10: Akkermans", if it is not a type, Be
Page 11 and 12: Straightness of the indument son1et
Page 13 and 14: Staminodia in the fourth whorl are
Page 15 and 16: elongs not to B. ova/is but to B. a
Page 17 and 18: Many species occur fron1 premontane
Page 19 and 20: Although I did not make an intensiv
Page 21 and 22: 12a. Inflorescences racen1ose, more
Page 23 and 24: 1. Beilscluniedia alloiophylla (Rus
Page 25 and 26: slopeS of creek, NE slope of Altos
Page 27 and 28: Rica. This group appears to have re
Page 29 and 30: anay, and he stated that the Colotn
Page 31 and 32: Phenology: collected in flower in M
Page 33 and 34: angustifolia has pubescent pistils,
Page 35 and 36: The specimen with a label saying "B
Page 37 and 38: 23, 1990 (fr.), Estinoza45 (CR, INB
Page 39 and 40: Beilschmiedia rohliana Lasser, Bol.
Page 41 and 42: (fl.), Haber 11837 (MO); La Peii.a
Page 43 and 44: (F); Con. Ceylan, path La Esmeralda
Page 45 and 46: Colombia (Cuatrecasas 22220, 22438)
Page 47 and 48: locality, Nov. 29- Dec. 18, 1964 (s
Page 49 and 50: Among Brazilian Beifschmiedia speci
Page 51 and 52: van der Werff et al. 12951 (holotyp
Page 53 and 54: inside; starnens 9, filaments ca. 0
Page 55 and 56: ecause the flowers of the type coll
Page 57 and 58: This species also shares a similar
Page 59: synonym of B . sulcata, supposedly
Page 63 and 64: Cristobal leaving main Hwy 100.8 mi
Page 65 and 66: pubescent with erect, curly to wavy
Page 67 and 68: Tree, to 20 m tall. Terminal buds p
Page 69 and 70: 16 x (5) 6-10 em; base obtuse to cu
Page 71 and 72: Alajuela: Canton de Alfaro Ruiz, Pa
Page 73 and 74: There are several collections from
Page 75 and 76: Spiritus (Las Villas): same mountai
Page 77 and 78: (fl.), Ekman Hl4565 (MO), Apr. 3, 1
Page 79 and 80: 316, 1906, Little Jr. 6293, Aguilar
Page 81 and 82: Cerro Hueco, ca. 700 malt., Feb. 4,
Page 83 and 84: ecause there is a distribution gap
Page 85 and 86: Beilschmiedia immersinervis also ap
Page 87 and 88: hairs. Fruits ellipsoid, ca. 5 x 3
Page 89 and 90: Additional specimens examined: Cost
Page 91 and 92: 2, 1994 (young fl.), van der Werff
Page 93 and 94: In the illustration of Ruiz and Pav
Page 95 and 96: Table 1. List of the Beilschmiedia
Page 97 and 98: Legends of figures Fig. 1. Five typ
Page 99: 4 A B c . w ; . -)1 . ..... J.f D f
Page 102 and 103: o Beilschmiedia berteroana * B. bre
Page 104 and 105: o Beilschmiedia hexanrhera * B. hon
Page 106 and 107: 0 o Beilschmiedia ovaLioides *B. ov
Page 108 and 109: C::::J.o•' :: ~ ·a ' \) ' ,o 0 .
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Chapter 3. Leaf Anatomy of neotropi
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3. Leaf sections The technique used
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16), B. ovalis (Fig. 18), B. peruiu
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smooth. However, in B. angustielfip
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uneven-sized subsidiary cells. This
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Midrib. The most distinct character
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This grouping is more or less suppo
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In leaf sectional study, neotropica
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Species of group 2 have leaves oppo
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Table 1. List of samples for venati
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0\ Table 2. Continued. B. emarginat
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Legends of figures Fig. 1-25. Venat
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Fig. 122-143. Midrib sections of ne
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2 132
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10 1 1 12 \ 14 134
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136
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140
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References Allen, C. K. 1945. Studi
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Meissner, C. F. 1864. Lauraceae. Pp
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