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Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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only from the type collections. And since Kostermans' s revision, 15 names have been<br />

already described under <strong>Beilschmiedia</strong> in the neotropics. A revision for <strong>neotropical</strong><br />

<strong>Beilschmiedia</strong> <strong>species</strong>, being up to date and with n1ore material, is needed for an<br />

understanding <strong>of</strong> the whole genus and also the <strong>neotropical</strong> <strong>Lauraceae</strong>.<br />

Fortunately, as Burger (1988) mentions, Latin American botanists are actively<br />

collecting <strong>neotropical</strong> plants including <strong>Lauraceae</strong> these days. Especially, van der Werff in the<br />

Missouri Botanical Garden has been working on a number <strong>of</strong> projects on a systematic <strong>study</strong><br />

<strong>of</strong> <strong>Lauraceae</strong> coupled with intensive collecting <strong>of</strong> the family in the neotropics. I had an<br />

opportunity to <strong>study</strong> at Missouri Botanical Garden from fall in 1994 to spring in 1997, and<br />

took the advantage to work on <strong>neotropical</strong> <strong>Beilschmiedia</strong> <strong>species</strong> under van der Werff's<br />

supervision.<br />

In this <strong>study</strong> I revised <strong>neotropical</strong> <strong>Beilschmiedia</strong> <strong>species</strong> in which I recognize 27<br />

<strong>species</strong>, with general descriptions <strong>of</strong> morphology, anatomy, palynology, karyology,<br />

distribution, reproductive biology, seed dispersal, and systematic position <strong>of</strong> the <strong>species</strong> in<br />

the fmnily in the second chapter. And for better understanding <strong>of</strong> relationships <strong>of</strong> the<br />

<strong>species</strong>, I employed leaf anatomy in the third chapter. Neotropical <strong>Beilschmiedia</strong> <strong>species</strong><br />

have relatively similar flowers and fruits except for a few <strong>species</strong>, and it is more reasonable<br />

to <strong>study</strong> vegetative n1orphology rather than reproductive morphology for systen1atics <strong>of</strong> the<br />

<strong>species</strong>. An1ong <strong>neotropical</strong> Beilscluniedia <strong>species</strong>, there appear to be two groups, one group<br />

<strong>of</strong> the <strong>species</strong> alternate leaves and fine ramification (ramification is here defined as ultimate<br />

branching patterns in leaf veins), and another group <strong>of</strong> the <strong>species</strong> with opposite leaves and<br />

coarse ratnification. Preliminary observation (Christophel, pers. comm.) indicated that the<br />

two groups also differ in characters <strong>of</strong> the cuticle. Therefore, I investigated leaf anaton1ical<br />

characters (venation patterns, cuticular and leaf section characters) for all the <strong>neotropical</strong><br />

Beilschrniedia <strong>species</strong> available in order to discuss the following; 1) whether the cuticular<br />

and leaf section characters support the grouping based on phyllotaxis and venation patterns,<br />

and 2) how we can incorporate leaf-anatomical characters in the classification <strong>of</strong> <strong>neotropical</strong><br />

<strong>Beilschmiedia</strong> <strong>species</strong>. These studies would not only help to know <strong>neotropical</strong> <strong>Beilschmiedia</strong><br />

<strong>species</strong> further, but would also give an important step to understand whole <strong>Beilschmiedia</strong>,<br />

one <strong>of</strong> the largest genera in <strong>Lauraceae</strong>.<br />

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