Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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2.5 x 1.5 em, the surface smooth; infructescence axis ca. 2 mm in diarn., fruit pedicels rarely slightly thickened to 3 mrn in diam., fruit pedicels slightly constricted at the base, or almost not constricted, without or with a few lenticels, slightly darker than the infructescence axis. Phenology: collected in flower in December and January, in mature fruit in March and June. Distribution: north-western part of Costa Rica, 300-1400 m alt. (Fig. 10). Habitat: premontane wet forest. Common name: unknown. Economic use: unknown. Additional specimens examined: Costa Rica. Alajuela: along a road between Canas and Upala, lower slopes Volcan Tenorio, 450 malt., Jan. 23, 1984 (fl.), Pennington 11416 (CR). Guanacaste: ParqueNacional Guanacaste, Estaci6n Pi till a, La Pasmompa, primary forest remnant beside the road to La Esperanza, 11 °02'N 8Y25.3'W, 300m alt., Jun. 17, 1989 (fr.),Hamme117502 (INB,MO); Monteverde zone, Canton deAbangares, Cordillera de Tilarcin, from Santa Elena to San Rafael, 10°20'00"N 84°53'00"W, 1200 malt., Jan. 7, 1992 (fl.), Guindon 36 (INB, MO); same zone, Los Tomos, Tilaran, Pacific side., 1300 malt., Apr. 14, 1987 (immature fr.), Haber 6967 (INB, F, MO); Canton de Tilaran, San Rafael to El Dos de Tilarcin, Pacific slope, 10°22'N 84°53'W, 800-1100 malt., Mar. 11, 1992 (fr.), Haberl1070 (INB, MO); San Rafael, 10°21 'N 84o52'W, ca. 1010 malt., Mar.10.1996 (immature fr.), Yasuda 1312 (INB, MO). Beilschmiedia immersinervis has alternate leaves and fine ramification. This species is distinguished from the other species with alternate leaves and fine ramification by a combination of the following three characters; short curly pubescence on the terminal buds and twigs, secondary veins immersed on both sides of the leaves and pubescent anthers. Some collections of B. immersinervis were identified as B. pendula, supposedly because B. immersinervis has a similar leaf shape as B. pendula. But one can separate B. immersinervis from B. pendula because B. immersinervis has erect and curly pubescence on the terminal buds while B. pendula has appressed and straight pubescence on the terminal buds. Beilschmiedia immersinervis has the same pubescence type and ramification pattern as B. steyermarkii, but B. immersinervis has pubescent anthers and tends to have firmly charta ceo us and symmetrical leaves, while B. steyermarkii has glabrous anthers and tends to have more coriaceous and asymmetrical leaves. 52
This species also shares a similar ramification pattern and pubescent anthers with B. riparia, but B. immersinervis is easily distinguished from B. riparia by its immersed secondary veins. Additionally, B. immersinervis differs from B. riparia by its having glaucous leaves and slightly longer flower pedicels, although there are a couple of exceptional collections with glaucous lower leaf surfaces in B. riparia. Beilschmiedia immersinervis also tends to have corky twigs which is not seen in B. riparia. 14. Beilschmiedia latifolia (Nees) Sa. Nishida comb. nov. Hufelandia latifolia Nees, Systema Laurinarum: 674. 1836. TYPE: Peru. locality unknown, 1835 (fl.), Matthew 1433 (ho1otype, E; isotypes, BM, E, K, LE not seen, OXF not seen). Tree, to 25 m tall. Terminal buds densely pubescent with brown to ferruginous, erect, long, wavy to straight hairs. Twigs terete, angular when young, densely pubescent with erect long hairs, less densely when older. Leaves alternate; petioles 0.5-1 (1.5) em long, flat above, pubescent with erect long hairs to glabrous, concolorous with twigs; blades coriaceous, ovate to broadly elliptic, (5) 7-11 (18) x 4-8 (11.5) em; base obtuse to round, not inrolled, apex obtuse; upper leaf surface glabrous, lower leaf surface densely pubescent with long hairs when young, less densely when old, rarely almost glabrous when old, lower surface glaucous; midrib immersed above, raised below, secondary veins 6-9 pairs, immersed or impressed, rarely slightly raised above, raised below, tertiary veins usually percurrent, rarely only weakly percurrent, ramification fine, areolate without free veinlets, immersed above, imn1ersed (the pattern visible) or slightly raised below. Inflorescences axillary, paniculate, 6-11 em long, densely pubescent with long erect hairs, with 20-40 flowers per inflorescence; flower pedicels of the lateral divisions 0.5-1 mm long, pedicels of the central flowers up to 2 mm long. Flowers (2.3) 2.5-3 mm long; tepals 6, equal, ovate, 1.7-2.2 mm long, 1-1.5 rnm wide; pubescent with erect hairs on both sides; stamens 9, outer six filaments ca. 0.3 mm long, innermost three filaments 0.3-0.5 mm long, filaments pubescent, anthers ca. 1 mm long, all the anthers 2-celled, apex of the anthers acute to truncate, apex of the anthers glabrous, glands of the innermost three stamens globose; stan1inodia 3, cordate in outline, 0.5-0.8 mm long; pistil ca. 1.4 mm long, glabrous, ovary as long as and gradually narrowed into the style; receptacle pubescent with appressed long hairs. Fruits ellipsoid, black, 3-5 x (1.5) 2 em, the surface smooth; infructescence axis 2-3 53
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KURENAI : Kyoto University Researc
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Contents Abstract Chapter 1 Chapter
Page 5 and 6: Chapter 1 General Introduction Laur
Page 7 and 8: Chapter 2 Revisional study of neotr
Page 9 and 10: Akkermans", if it is not a type, Be
Page 11 and 12: Straightness of the indument son1et
Page 13 and 14: Staminodia in the fourth whorl are
Page 15 and 16: elongs not to B. ova/is but to B. a
Page 17 and 18: Many species occur fron1 premontane
Page 19 and 20: Although I did not make an intensiv
Page 21 and 22: 12a. Inflorescences racen1ose, more
Page 23 and 24: 1. Beilscluniedia alloiophylla (Rus
Page 25 and 26: slopeS of creek, NE slope of Altos
Page 27 and 28: Rica. This group appears to have re
Page 29 and 30: anay, and he stated that the Colotn
Page 31 and 32: Phenology: collected in flower in M
Page 33 and 34: angustifolia has pubescent pistils,
Page 35 and 36: The specimen with a label saying "B
Page 37 and 38: 23, 1990 (fr.), Estinoza45 (CR, INB
Page 39 and 40: Beilschmiedia rohliana Lasser, Bol.
Page 41 and 42: (fl.), Haber 11837 (MO); La Peii.a
Page 43 and 44: (F); Con. Ceylan, path La Esmeralda
Page 45 and 46: Colombia (Cuatrecasas 22220, 22438)
Page 47 and 48: locality, Nov. 29- Dec. 18, 1964 (s
Page 49 and 50: Among Brazilian Beifschmiedia speci
Page 51 and 52: van der Werff et al. 12951 (holotyp
Page 53 and 54: inside; starnens 9, filaments ca. 0
Page 55: ecause the flowers of the type coll
Page 59 and 60: synonym of B . sulcata, supposedly
Page 61 and 62: Additional specimen examined: Brazi
Page 63 and 64: Cristobal leaving main Hwy 100.8 mi
Page 65 and 66: pubescent with erect, curly to wavy
Page 67 and 68: Tree, to 20 m tall. Terminal buds p
Page 69 and 70: 16 x (5) 6-10 em; base obtuse to cu
Page 71 and 72: Alajuela: Canton de Alfaro Ruiz, Pa
Page 73 and 74: There are several collections from
Page 75 and 76: Spiritus (Las Villas): same mountai
Page 77 and 78: (fl.), Ekman Hl4565 (MO), Apr. 3, 1
Page 79 and 80: 316, 1906, Little Jr. 6293, Aguilar
Page 81 and 82: Cerro Hueco, ca. 700 malt., Feb. 4,
Page 83 and 84: ecause there is a distribution gap
Page 85 and 86: Beilschmiedia immersinervis also ap
Page 87 and 88: hairs. Fruits ellipsoid, ca. 5 x 3
Page 89 and 90: Additional specimens examined: Cost
Page 91 and 92: 2, 1994 (young fl.), van der Werff
Page 93 and 94: In the illustration of Ruiz and Pav
Page 95 and 96: Table 1. List of the Beilschmiedia
Page 97 and 98: Legends of figures Fig. 1. Five typ
Page 99: 4 A B c . w ; . -)1 . ..... J.f D f
Page 102 and 103: o Beilschmiedia berteroana * B. bre
Page 104 and 105: o Beilschmiedia hexanrhera * B. hon
Page 106 and 107:
0 o Beilschmiedia ovaLioides *B. ov
Page 108 and 109:
C::::J.o•' :: ~ ·a ' \) ' ,o 0 .
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Chapter 3. Leaf Anatomy of neotropi
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3. Leaf sections The technique used
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16), B. ovalis (Fig. 18), B. peruiu
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smooth. However, in B. angustielfip
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uneven-sized subsidiary cells. This
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Midrib. The most distinct character
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This grouping is more or less suppo
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In leaf sectional study, neotropica
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Species of group 2 have leaves oppo
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Table 1. List of samples for venati
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0\ Table 2. Continued. B. emarginat
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Legends of figures Fig. 1-25. Venat
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Fig. 122-143. Midrib sections of ne
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2 132
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10 1 1 12 \ 14 134
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136
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138
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140
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142
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References Allen, C. K. 1945. Studi
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Meissner, C. F. 1864. Lauraceae. Pp
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