Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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emarginata(Meissner) Mez, Jahrb. Konig!. Bot. Gart. Berl. 5: 18. 1889. TYPE: Brazil. Sao Paulo: near Lorena, Oct., 1833 (fl.), Riede/1585 (holotype, LE; isotypes, G-BOIS not seen, K, NY not seen). Tree, to 15 n1 tall. Tem1inal buds pubescent with rusty erect, short curly hairs. Twigs terete, compressed when young, sparsely pubescent with erect short curly hairs or almost glabrous. Leaves opposite; petioles ca. 0.5 em long, flat above, sparsely pubescent or glabrous, concolorous with or slightly darker than twigs; blades chartaceous, elliptic, often asymmetrical and sinuate, 6.5-12 x 3-6.5 cn1; base cuneate, not inrolled, apex emarginate to round; leaf surface glabrous on both sides, uncertain whether lower leaf surface glaucous or not; n1idrib immersed above, raised below, secondary veins 7-8 pairs, slightly raised or almost in1mersed above, raised below, tertiary veins not percurrent, ramification coarse, square-like with free veinlets, slightly raised or almost in1mersed above, slightly raised below. Inflorescences axillary, paniculate, 3-5 em long, aln1ost glabrous or sparsely pubescent with erect short curly hairs, with 15-30 flowers per inflorescence; flower pedicels of the lateral divisions 0.5-1.2 mm long, pedicels of the central flowers up to 3 n1mlong. Flowers 2-2.8 mm long, 2.2-2.8 mm in diam.; tepals 6, equal, ovate, 1.2-1.5 mm long, 0.8-1.2 mm wide, almost glabrous outside, sparsely pubescent with appressed hairs inside; stan1ens 9, outer six filaments ca. 0.2 1111n long, innermost three filaments ca. 0.5 mm long, filaments pubescent, outer six anthers ca. 0.9 mm long, innermost three anthers ca. 0.7 mn1 long, all the anthers 2-celled, apex of the anthers obtuse to truncate, anthers pubescent, glands of innermost three stamens globose; stan1inodia 3, ca. 0.7 mm long; pistil 1-1.5 mm long, glabrous, ovary as long as pistil and gradually narrowed into the style; receptacle pubescent with appressed hairs. Fruits unknown. Phenology: collected in flower in October and December. Distribution: Sao Paulo, Brazil (Fig. 9). Habitat: unknown. Common nan1e: unknown. Economic use: unknown. Additional specimen examined: Brazil. Sao Paulo: native in Botanical Garden, Dec. 9, 1955 (sterile), Kuhlmann 3885 (NY). 44
Among Brazilian Beifschmiedia species, B. emnrginata is distinguished by its rusty, short and erect pubescence on the tem1inal buds and twigs. Additionally, this species usually has undulate leaves and emarginate leaf apices, both of which are not seen in the other Brazilian species. Two collections by Hatschbach (2455 &1631 0) from Parana approach to B. emnrginata in vegetative characters, but have glabrous anthers. Species with pubescent anthers can rarely have exceptional collections which lack the pubescence, so these Hatschbach collections might still belong to B. emnrginata. But I decided to keep them separated from B. emarginata as indetem1inate, because the locality and the phenology are different from the other collections (the collections fron1 Parana were in flower in August and September). Actually, I have seen only two collections which can be identified as B. emnrginata. More material is needed to clearly delimit this species. 10. Beilschnziedia flunzinensis Kostermans, Rec. Trav. Bot. Neerl. 35: 865. 1938. TYPE: Brazil. Rio de Janeiro: Estado Rio G. Portella, Monte Sinai, 1935 (fl.), Nunes 323 (holotype, U not seen; isotypes, RB-2 sheets). Tree, to 15m tall. Tem1inal buds pubescent with appressed straight hairs. Twigs terete, slightly con1pressed when young, almost glabrous or sparsely pubescent with appressed hairs. Leaves opposite; petioles 1-2 em long, canaliculate above, glabrous, slightly darker than twigs; blades finnly chartaceous, elliptic, (1 0) 13-16 (18) x 4-7 em; base cuneate, slightly inrolled, apex acute, rarely obtuse; leaf surface glabrous on both sides, rarely lower leaf surface sparsely pubescent with short appressed hairs, lower leaf surface not glaucous; midrib immersed above, raised below, secondary veins 12-14 pairs, raised on both sides, tertiary veins not percurrent, ramification coarse, square-like with free veinlets, slightly raised above, raised below. Inflorescences axillary, paniculate, 4-7 cn1 long, glabrous to sparsely pubescent with erect hairs, with 20-35 flowers per inflorescence; flower pedicels of the lateral divisions ca. 1 111111 long, pedicels of the central flowers up to 2.5 mm long. Flowers greenish, 1.7-2 mm long, 2-2.5 mm in dian1.; tepals 6, equal, elliptic to ovate, 1- 1.5 mn1long, 1.1-1.5 mn1 wide, glabrous outside, sparsely pubescent with appressed hairs inside; stamens 9, outer six filaments ca. 0.2 mm long, innermost three filaments ca. 0.4 mm long, filaments pubescent, anthers 0.7-0.9 nun long, all the anthers 2-celled, apex of the 45
Page 1 and 2: KURENAI : Kyoto University Researc
Page 3 and 4: Contents Abstract Chapter 1 Chapter
Page 5 and 6: Chapter 1 General Introduction Laur
Page 7 and 8: Chapter 2 Revisional study of neotr
Page 9 and 10: Akkermans", if it is not a type, Be
Page 11 and 12: Straightness of the indument son1et
Page 13 and 14: Staminodia in the fourth whorl are
Page 15 and 16: elongs not to B. ova/is but to B. a
Page 17 and 18: Many species occur fron1 premontane
Page 19 and 20: Although I did not make an intensiv
Page 21 and 22: 12a. Inflorescences racen1ose, more
Page 23 and 24: 1. Beilscluniedia alloiophylla (Rus
Page 25 and 26: slopeS of creek, NE slope of Altos
Page 27 and 28: Rica. This group appears to have re
Page 29 and 30: anay, and he stated that the Colotn
Page 31 and 32: Phenology: collected in flower in M
Page 33 and 34: angustifolia has pubescent pistils,
Page 35 and 36: The specimen with a label saying "B
Page 37 and 38: 23, 1990 (fr.), Estinoza45 (CR, INB
Page 39 and 40: Beilschmiedia rohliana Lasser, Bol.
Page 41 and 42: (fl.), Haber 11837 (MO); La Peii.a
Page 43 and 44: (F); Con. Ceylan, path La Esmeralda
Page 45 and 46: Colombia (Cuatrecasas 22220, 22438)
Page 47: locality, Nov. 29- Dec. 18, 1964 (s
Page 51 and 52: van der Werff et al. 12951 (holotyp
Page 53 and 54: inside; starnens 9, filaments ca. 0
Page 55 and 56: ecause the flowers of the type coll
Page 57 and 58: This species also shares a similar
Page 59 and 60: synonym of B . sulcata, supposedly
Page 61 and 62: Additional specimen examined: Brazi
Page 63 and 64: Cristobal leaving main Hwy 100.8 mi
Page 65 and 66: pubescent with erect, curly to wavy
Page 67 and 68: Tree, to 20 m tall. Terminal buds p
Page 69 and 70: 16 x (5) 6-10 em; base obtuse to cu
Page 71 and 72: Alajuela: Canton de Alfaro Ruiz, Pa
Page 73 and 74: There are several collections from
Page 75 and 76: Spiritus (Las Villas): same mountai
Page 77 and 78: (fl.), Ekman Hl4565 (MO), Apr. 3, 1
Page 79 and 80: 316, 1906, Little Jr. 6293, Aguilar
Page 81 and 82: Cerro Hueco, ca. 700 malt., Feb. 4,
Page 83 and 84: ecause there is a distribution gap
Page 85 and 86: Beilschmiedia immersinervis also ap
Page 87 and 88: hairs. Fruits ellipsoid, ca. 5 x 3
Page 89 and 90: Additional specimens examined: Cost
Page 91 and 92: 2, 1994 (young fl.), van der Werff
Page 93 and 94: In the illustration of Ruiz and Pav
Page 95 and 96: Table 1. List of the Beilschmiedia
Page 97 and 98: Legends of figures Fig. 1. Five typ
Page 99:
4 A B c . w ; . -)1 . ..... J.f D f
Page 102 and 103:
o Beilschmiedia berteroana * B. bre
Page 104 and 105:
o Beilschmiedia hexanrhera * B. hon
Page 106 and 107:
0 o Beilschmiedia ovaLioides *B. ov
Page 108 and 109:
C::::J.o•' :: ~ ·a ' \) ' ,o 0 .
Page 110 and 111:
Chapter 3. Leaf Anatomy of neotropi
Page 112 and 113:
3. Leaf sections The technique used
Page 114 and 115:
16), B. ovalis (Fig. 18), B. peruiu
Page 116 and 117:
smooth. However, in B. angustielfip
Page 118 and 119:
uneven-sized subsidiary cells. This
Page 120 and 121:
Midrib. The most distinct character
Page 122 and 123:
This grouping is more or less suppo
Page 124 and 125:
In leaf sectional study, neotropica
Page 126 and 127:
Species of group 2 have leaves oppo
Page 128 and 129:
Table 1. List of samples for venati
Page 130 and 131:
0\ Table 2. Continued. B. emarginat
Page 132 and 133:
Legends of figures Fig. 1-25. Venat
Page 134 and 135:
Fig. 122-143. Midrib sections of ne
Page 136 and 137:
2 132
Page 138 and 139:
10 1 1 12 \ 14 134
Page 140 and 141:
136
Page 142 and 143:
138
Page 144 and 145:
140
Page 146 and 147:
142
Page 148 and 149:
144
Page 150 and 151:
146
Page 152 and 153:
148
Page 154 and 155:
150
Page 156 and 157:
152
Page 158 and 159:
154
Page 160 and 161:
References Allen, C. K. 1945. Studi
Page 162:
Meissner, C. F. 1864. Lauraceae. Pp
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