Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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Key to neotropical Beilschntiedia species. 1 a. Leaves strictly opposite. Ramification on lower leaf surface coarse, smallest meshes (mesh is defined here as a delicate network of a minor venation) larger than 1.5 mm in diam. 2a. Apex of anthers glabrous. Fron1 Central America or Chile. 3a. Leaves roundish-ovate. Leaf apex rounded to retuse. Frmn Chile. 4a. Tepals glabrous outside. Indument on the terminal buds and twigs appressed or spreading, straight. . .. ..... . ... . .. ... ... .. . .. ... . ... . ... ... .... .. ... ... .. ..... B. berteroana. 4 b. Tepals pubescent outside. Indument on the terminal buds and twigs erect, curly to wavy. ... ...... .. ..... ............ .......... . ........................ ........ .. ... ... B. miersii. 3 b. Leaves elliptic. Leaf apex acute or acuminate. From Central America. Sa. Secondary veins more than 14 pairs. Indument on the twigs erect, long and wavy . ... .. ... . .. ... . . ... .... ... ....... .... .. ... . ... . ... .... ... ... .... ..... .. .. ... . . B. angustieLLiptica. Sb. Secondary veins less than 13 pairs. Indmnent on the twigs appressed or spreading, or twigs glabrous. 6a. Flower pedicels shorter than 1.5 n1n1long ....... . ............. ..... .... B . hondurensis. 6b. Flower pedicels longer than 3.0 mm long. ...................... .. ...... .. .. B. brenesii. 2 b. Apex of anthers pubescent. Fr01n northern South An1erica or Brazil. 7 a. Lower leaf surface pubescent with erect, long and straight hairs. Leaf base obtuse. B. taubertiana. 7 b. Lower leaf surface glabrous. Leaf base cuneate. 8a. Indun1ent on the terminal buds and twigs rusty, erect, very short and curly. B. emarginata. 8 b. Indument on the ten11inal buds and twigs not rusty, appressed, relatively long and straight. 9a. Leaves obovate. Leaf apex round or roundish-obtuse. 10 a . Flowers longer than 3 n1m, wider than 3 111m. Flowers globose. .. .. .. .. B. rig ida. 10 b. Flowers shorter than 2.5 mn1, narrower than 2. 7 111111. Flowers depressedglobose. ....... .. ... ........ ............... ............... ........ ...... B. Linharesensis. 9 b. Leaves elliptic. Leaf apex acute (rarely obtuse). lla. Leaf width less than 1/3 of the leaf length . .. ...... .. ........ .. ....... B. angustifoLia. 11 b. Leaf width more than 1/2 of the leaf length. 16
12a. Inflorescences racen1ose, more or less densely pubescent. ............... B. stricta. 12 b. Inflorescences paniculate, sparsely pubescent or almost glabrous. 13a. Leaf base slightly inrolled. From Brazil............................ B . fluminensis. 13 b. Leaf base flat. From northern South America. . . . . . . . . . . . . . . . . . . . . . B. curviramea. 1 b. Leaves alternate, rarely sub-opposite. Ran1ification usually fine, smallest n1eshes sn1aller than 0.5 mm in diam. When ramification coarse, leaves still alternate and from Ecuador or Colombia (B. costaricensis). 14a. Fertile stamens 6. Staminodia 6. Fron1 French Guiana. .. . . . . .. .. . . .. . . . . B. hexanthera. 14 b. Fertile stamens 9. Statninodia 3. Fron1 Central America, West Indies, or western South An1erica. lSa. Indument of the tern1inal buds and twigs appressed or spreading (when indument spreading, ran1ification slightly less fine, meshes around 0.5 mm in diam., with free vein lets). 16 a. Lower leaf surface glabrous on tissue or very rarely pubescent with spreading hairs only along the major veins (when pubescent, ramification square-like with free veinlets). 17 a. Ramification with few free veinlets. From Mexico. . . . . . . . . . . . . . . . . . . . . . . B . mexicana 17b. Ramification with free veinlets. From Central America north to Nicaragua, West Indies, or South America. 18a Ramification immersed above (although the pattern visible), almost immersed or slightly raised below. Lower leaf surface glaucous. Fruit pedicels constricted at the base. . .. .... ............................................................... B . pendula. 18b. Ramification raised above, slightly to conspicuously raised below. Lower leaf surface not glaucous. Fruit pedicels not constricted at the base. B. costaricensis. 16 b. Lower leaf surface pubescent with appressed hairs on tissue. Ran1ification with few free veinlets. 19a. Ramification square-like. Ramification pattern always only partially visible above. Ramification pattern visible above much coarser than the ramification pattern below. .. .. ... .... .............................................................. B. pendula. 19 b. Ramification finely areolate. Ran1ification pattern above completely invisible or slightly visible. When ramification pattern visible above, even the finest pattern visible. ...... ..... .. ........ ................................................. B. tovarensis. 17
Page 1 and 2: KURENAI : Kyoto University Researc
Page 3 and 4: Contents Abstract Chapter 1 Chapter
Page 5 and 6: Chapter 1 General Introduction Laur
Page 7 and 8: Chapter 2 Revisional study of neotr
Page 9 and 10: Akkermans", if it is not a type, Be
Page 11 and 12: Straightness of the indument son1et
Page 13 and 14: Staminodia in the fourth whorl are
Page 15 and 16: elongs not to B. ova/is but to B. a
Page 17 and 18: Many species occur fron1 premontane
Page 19: Although I did not make an intensiv
Page 23 and 24: 1. Beilscluniedia alloiophylla (Rus
Page 25 and 26: slopeS of creek, NE slope of Altos
Page 27 and 28: Rica. This group appears to have re
Page 29 and 30: anay, and he stated that the Colotn
Page 31 and 32: Phenology: collected in flower in M
Page 33 and 34: angustifolia has pubescent pistils,
Page 35 and 36: The specimen with a label saying "B
Page 37 and 38: 23, 1990 (fr.), Estinoza45 (CR, INB
Page 39 and 40: Beilschmiedia rohliana Lasser, Bol.
Page 41 and 42: (fl.), Haber 11837 (MO); La Peii.a
Page 43 and 44: (F); Con. Ceylan, path La Esmeralda
Page 45 and 46: Colombia (Cuatrecasas 22220, 22438)
Page 47 and 48: locality, Nov. 29- Dec. 18, 1964 (s
Page 49 and 50: Among Brazilian Beifschmiedia speci
Page 51 and 52: van der Werff et al. 12951 (holotyp
Page 53 and 54: inside; starnens 9, filaments ca. 0
Page 55 and 56: ecause the flowers of the type coll
Page 57 and 58: This species also shares a similar
Page 59 and 60: synonym of B . sulcata, supposedly
Page 61 and 62: Additional specimen examined: Brazi
Page 63 and 64: Cristobal leaving main Hwy 100.8 mi
Page 65 and 66: pubescent with erect, curly to wavy
Page 67 and 68: Tree, to 20 m tall. Terminal buds p
Page 69 and 70: 16 x (5) 6-10 em; base obtuse to cu
Page 71 and 72:
Alajuela: Canton de Alfaro Ruiz, Pa
Page 73 and 74:
There are several collections from
Page 75 and 76:
Spiritus (Las Villas): same mountai
Page 77 and 78:
(fl.), Ekman Hl4565 (MO), Apr. 3, 1
Page 79 and 80:
316, 1906, Little Jr. 6293, Aguilar
Page 81 and 82:
Cerro Hueco, ca. 700 malt., Feb. 4,
Page 83 and 84:
ecause there is a distribution gap
Page 85 and 86:
Beilschmiedia immersinervis also ap
Page 87 and 88:
hairs. Fruits ellipsoid, ca. 5 x 3
Page 89 and 90:
Additional specimens examined: Cost
Page 91 and 92:
2, 1994 (young fl.), van der Werff
Page 93 and 94:
In the illustration of Ruiz and Pav
Page 95 and 96:
Table 1. List of the Beilschmiedia
Page 97 and 98:
Legends of figures Fig. 1. Five typ
Page 99:
4 A B c . w ; . -)1 . ..... J.f D f
Page 102 and 103:
o Beilschmiedia berteroana * B. bre
Page 104 and 105:
o Beilschmiedia hexanrhera * B. hon
Page 106 and 107:
0 o Beilschmiedia ovaLioides *B. ov
Page 108 and 109:
C::::J.o•' :: ~ ·a ' \) ' ,o 0 .
Page 110 and 111:
Chapter 3. Leaf Anatomy of neotropi
Page 112 and 113:
3. Leaf sections The technique used
Page 114 and 115:
16), B. ovalis (Fig. 18), B. peruiu
Page 116 and 117:
smooth. However, in B. angustielfip
Page 118 and 119:
uneven-sized subsidiary cells. This
Page 120 and 121:
Midrib. The most distinct character
Page 122 and 123:
This grouping is more or less suppo
Page 124 and 125:
In leaf sectional study, neotropica
Page 126 and 127:
Species of group 2 have leaves oppo
Page 128 and 129:
Table 1. List of samples for venati
Page 130 and 131:
0\ Table 2. Continued. B. emarginat
Page 132 and 133:
Legends of figures Fig. 1-25. Venat
Page 134 and 135:
Fig. 122-143. Midrib sections of ne
Page 136 and 137:
2 132
Page 138 and 139:
10 1 1 12 \ 14 134
Page 140 and 141:
136
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138
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140
Page 146 and 147:
142
Page 148 and 149:
144
Page 150 and 151:
146
Page 152 and 153:
148
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150
Page 156 and 157:
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154
Page 160 and 161:
References Allen, C. K. 1945. Studi
Page 162:
Meissner, C. F. 1864. Lauraceae. Pp
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Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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