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Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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spinules, exine is ectexinous, intine is a dominant layer <strong>of</strong> the pollen wall and uniformly<br />

thick, possibly equally suited for pollen tube egress.<br />

But these <strong>species</strong> (and Mezilaurus) are different from the other genera in size <strong>of</strong><br />

spinules and in surface <strong>of</strong> the basal cushions and exine. Size <strong>of</strong> the spinules is relatively<br />

large, 1-1.5 j.lm long. Surface <strong>of</strong> the basal cushions and exine is without granular processes.<br />

Grain size <strong>of</strong> B. 1niersii is reported as the largest <strong>of</strong> all the <strong>species</strong> they studied. Density<br />

<strong>of</strong> spinules is reported as one <strong>of</strong> the lowest in their <strong>study</strong>.<br />

Karyology<br />

Among <strong>neotropical</strong> <strong>Beilschmiedia</strong> <strong>species</strong>, only B. berteroana has been reported in<br />

chromosome number. The chromoson1e number is x = 12 (2n = 24) (Raven 1975), which<br />

is con1mon in the <strong>Lauraceae</strong> <strong>species</strong>. Polyploid numbers have not been found in<br />

<strong>Beilschmiedia</strong>.<br />

Distribution<br />

<strong>Beilschmiedia</strong> <strong>species</strong> distributed in the <strong>neotropical</strong> counties are listed in Table 1.<br />

In the neotropics, <strong>Beilschmiedia</strong> ranges from central Mexico to southern Brazil and<br />

Chile. <strong>Beilschmiedia</strong> is absent from An1azon basin and northern Chile, therefore the<br />

distribution appears to consist <strong>of</strong> one large region from central Mexico to western Bolivia,<br />

and sn1all regions in northern South America, south-eastern Brazil and central Chile (Fig. 6).<br />

In north-western Latin America, distribution <strong>of</strong> <strong>Beilschmiedia</strong> ranges from central<br />

Mexico to western Bolivia (Fig. 6 A). The country with the highest number <strong>of</strong> <strong>species</strong> is<br />

Costa Rica, where eight <strong>species</strong> occur. Six <strong>of</strong> these <strong>species</strong> have also represent from<br />

Panarna, and four <strong>of</strong> them are also distributed from Venezuela to Ecuador. On the other<br />

hand, only two <strong>of</strong> the Costarican <strong>species</strong>, B. ovalis and B. pendula, extend the distribution<br />

north <strong>of</strong> Costa Rica (to Honduras). <strong>Beilschmiedia</strong> pendula is the <strong>species</strong> with the widest<br />

distribution in <strong>neotropical</strong> Beilsch1niedia <strong>species</strong>, which occurs from Honduras to Ecuador,<br />

including the West Indies. In Mexico there are six <strong>species</strong> present, four <strong>of</strong> which are<br />

enden1ic to Mexico. In the Guianas, there are two <strong>species</strong> distributed; B. curviramea is<br />

known from eastern Venezuela to Guyana (Fig. 6 B), and B. hexanthera is known only from<br />

French Guiana (Fig. 6 C).<br />

12

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