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Revisional study of neotropical Beilschmiedia species (Lauraceae ...

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uneven-sized subsidiary cells. This is also true <strong>of</strong> <strong>neotropical</strong> <strong>Beilschmiedia</strong> <strong>species</strong>. Distinct<br />

subsidiary cells, whose periclinal walls are irregularly thickened and appear granular, occur<br />

in B. angustifolia (Fig. 100), B. curviramea (Fig. 104), B. linharesensis (Fig. 109) and B.<br />

rigida (Fig. 116).<br />

7) The nature <strong>of</strong> the stomatal/edges. Many <strong>neotropical</strong> <strong>Beilschmiedia</strong> <strong>species</strong> have narrow<br />

stomatal ledges. Slightly wider stomatal ledges are seen in B. angustifolia (Fig. 100), B.<br />

curviramea (Fig. 104), B. linharesensis (Fig. 109) and B. rigida (Fig. 116). Extremely wide<br />

stomatal ledges occur in B. angustielliptica (Fig. 99), B. brenesii (Fig. 102), B. hondurensis<br />

(Fig. 106), whose inner stomatal ledges protrude between the guard cells and the subsidiary<br />

cells, forming "butterfly-shape" (termed in Christophel et al. 1996). B. berteroana also has<br />

very different stomata from the other <strong>species</strong>, in which the inner stomatal ledges protrude in<br />

parallel to the outer ledges, forming rectangular "box-shape" (termed by Christophel, pers.<br />

comm.) and the ledges themselves are scarcely stained (Fig. 101).<br />

8) Presence/absence and chnracters <strong>of</strong> various specialized cells. The specialized cells mostly<br />

seen in <strong>neotropical</strong> <strong>Beilschmiedia</strong> <strong>species</strong> are trichomes, which are same as all the trichomes<br />

<strong>of</strong> <strong>Lauraceae</strong> thus far reported (Christophel et al. 1996). The trichomes <strong>of</strong> <strong>neotropical</strong><br />

<strong>Beilschmiedia</strong> <strong>species</strong> are simple, with their poral bases (e.g., Fig. 86). Although the<br />

trichomes themselves are various among the <strong>species</strong> (see the section indument in the second<br />

chapter), basal cells <strong>of</strong> trichomes are relatively uniform. Giant stomata, referred as<br />

"hydathodes" in Christophel et. al. (1996), are sometimes present in <strong>neotropical</strong><br />

<strong>Beilschmiedia</strong> <strong>species</strong> (e.g., Fig. 97), but there is no significant difference among the<br />

<strong>species</strong>.<br />

3. Leaf sections<br />

Twenty-two <strong>of</strong> 27 <strong>species</strong> were examined for the leaf section <strong>study</strong>. <strong>Beilschmiedia</strong><br />

anay, B . emarginata, B. fluminensis, B. stricta and B . taubertiana could not be studied<br />

because <strong>of</strong> the lack <strong>of</strong> material. Pictures <strong>of</strong> the petiole sections are shown in Fig. 120-121,<br />

midrib sections in Fig. 122-143.<br />

Hypodermis. According to Metcalfe (1987), a single-layered hypodermis has been recorded<br />

beneath the adaxial epidermis in <strong>species</strong> <strong>of</strong> <strong>Beilschmiedia</strong>.<br />

114

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