Revisional study of neotropical Beilschmiedia species (Lauraceae ...

pseudo-brochidodromous (in which the leaf appears brochidodromous but actually the loops
are formed by strengthened tertiaries, not by the secondaries themselves).
Most neotropical Beilschmiedia species have pseudo-brochidodromous leaves. But the
leaves of these species sometimes have the secondaries near the leaf apex, forming loops by
themselves and appearing partly brochidodromous. Leaves of B. angustifolia (Fig. 4) andB.
linharesensis (Fig. 14) in the X-ray pictures appear to be entirely brochidodromous, but the
larger leaves of these species often have pseudo-brochidodromous (and partly
brochidodromous). A few species such as B. ova/is (Fig. 18) and B. steyermarikii (Fig. 22)
appear to have eucamptodromous leaves, but even in these species margins of the
secondaries curve inwards and some of the tertiaries reach to the adjust veins, appearing
pseudo-brochidodromous.
Tertiary vein pattern. In the tertiaries, Christophel and Rowett (1996) recognized two
patterns; 1) percurrent, in which the tertiaries directly connect to adjacent secondaries, 2)
reticulate, in which tertiaries anastomose with other tertiary or secondary veins and do not
run directly to secondaries. The former category, percurrent, can be divided into two further
categories; 1) strongly percurrent, having a ladder-like appearance, 2) weakly percurrent,
appearing to meander or arch in their course. The latter category can be also divided into two
further categories, 1) orthogonal, meaning that they form mainly right-angled anastomoses,
2) random, meaning they are either curved or form a variety of anastomoses.
Some neotropical Beilschmiedia species, such as B . alloiophylla (Fig. 1), B. anay (Fig.
2), B. latifolia (Fig. 13), B. obovatifolia (Fig. 17), B. ovalioides, and B. riparia (Fig. 21),
have strongly percurrent tertiaries. Most of these species have mostly simple tertiaries, but
B . latifolia and B. riparia often have branched tertiaries. Weakly percurrent tertiaries often
occur in B. costaricensis (Fig. 7), B. hexanthera (Fig. 10), B. immersinervis (Fig. 12), B.
ova/is (Fig. 18), B. steyermarkii (Fig. 22) and B. tovarensis (Fig. 25). But B. costaricensis,
B. hexanthera, B. immersinervis and B. ova/is sometimes have reticulate rather than
percurrent tertiaries. The tertiaries in these species are mostly branched even if percurrent.
Reticulate tertiaries occur in B. angustielliptica (Fig. 3), B . angustifolia (Fig. 4), B.
berteroana (Fig. 5), B. brenesii (Fig. 6), B. costaricensis (Fig. 7),B. curviramea(Fig. 8),B.
emarginata, B. fluminensis (Fig. 9), B. hexanthera (Fig. 10), B. hondurensis (Fig. 11), B.
immersinervis (Fig. 12), B. linharesensis (Fig. 14), B. mexicana (Fig. 15), B. miersii (Fig.
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