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Hayes and Garber - Cucurbit Breeding

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6 BREEDING CROP PLANTS<br />

more attention to the planting of their crops so that those of like<br />

kind did not grow so near each other that crossing through the<br />

aid of insects would take place.<br />

Sprengel, in a book published in<br />

1793, showed the important role played by insects in pollination<br />

<strong>and</strong> studied the adaptations for crossing found in many flowers.<br />

He concluded that nature intended flowers should not be pollinated<br />

by their own pollen.<br />

The Great Hybridist Gartner. In extent <strong>and</strong> number of his<br />

experiments Gartner's work is very great. In 1835 he heard of<br />

the offer of a prize made by the Dutch Academy of Sciences at<br />

Haarlem regarding the place of hybridization in producing<br />

new varieties of economic <strong>and</strong> ornamental plants.<br />

Gartner's paper on this question, which received the prize, was<br />

published in extended form in 1849. He made thous<strong>and</strong>s<br />

of crosses, involving nearly 700 species, <strong>and</strong> obtained about 250<br />

hybrids. The work was so carefully controlled <strong>and</strong> checked<br />

that the fact of sex in plants was thoroughly proved. He<br />

made a classification of hybrids according to whether they<br />

resembled one or the other parent in all respects, whether they<br />

resembled one parent in one part of the plant <strong>and</strong> the other<br />

parent in some other characters, or whether there was an almost<br />

equal balance. In the last case in later generations, the inclination<br />

toward the one or the other parent was supposed to be<br />

due to a slight overbalance of one or the other of the fertilizing<br />

materials. Gartner explains the appearance of the first hybrid<br />

generation as due to an inner force operating according to law.<br />

He, like Koelreuter <strong>and</strong> Weigmann, observed increased vigor in<br />

hybrids.<br />

He made experiments to determine the immediate effect of<br />

pollen with crosses between colorless <strong>and</strong> colored pericarp<br />

varieties of maize <strong>and</strong> in crosses between a brown-seeded Lychnis<br />

<strong>and</strong> one with a gray seed. As no change occurred, a law was<br />

developed to the effect that pollen does not immediately affect<br />

forms <strong>and</strong> external characters of seeds but influences the development<br />

of the resultant plant. He observed an immediate effect<br />

in some pea crosses <strong>and</strong> learned that the yellow cotyledon color<br />

dominated the green in the hybrid seeds.<br />

Early English Plant Breeders. Knight, Goss, <strong>and</strong> Herbert,<br />

three English workers, did much to develop the art of breeding.<br />

Knight, who was a practical horticulturist, recognized the aid<br />

of artificial cross-pollination in producing new kinds. He

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