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Hayes and Garber - Cucurbit Breeding

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268 BREEDING CROP PLANTS<br />

condition of most fruit material.<br />

A commercial variety may be<br />

extremely valuable <strong>and</strong> yet be heterozygous for many characters.<br />

On the other h<strong>and</strong>, the commercial variety may be homozygous<br />

for a large part of its characters. It seems reasonable to conclude<br />

that the more nearly homozygous the parental variety,<br />

other things being equal, the greater value it would have as a<br />

parent.<br />

The ability of impressing its characteristics upon the larger<br />

part of its offspring has been called prepotency by animal breeders.<br />

Such prepotency is genetically explained by the supposition that<br />

the prepotent parent is homozygous for certain dominant factors<br />

for the characters under observation. Hedrick <strong>and</strong> Wellington<br />

(1912) showed that some crosses between apple varieties produced<br />

a considerable percentage of individuals with small fruits.<br />

Thus the cross between Rails <strong>and</strong> Northern Spy gave great<br />

variability in size of apples, while the cross between Sutton <strong>and</strong><br />

Northern Spy gave progeny in which no trees were obtained<br />

which produced small fruit. One of the great difficulties is that<br />

it takes several years to learn the varieties which when crossed<br />

will give certain desired combination.<br />

Another difficulty which must be considered is that many<br />

varieties of fruits are self-sterile. This is of utmost importance<br />

in commercial fruit production for it is necessary to interplant<br />

such a variety with some variety which produces an abundance of<br />

pollen which is capable of fertilizing the variety in question <strong>and</strong><br />

TABLE LXXI.<br />

INCREASE IN WEIGHT OF SEED AND FRUIT DUE TO CROSS-<br />

POLLINATION<br />

Pollination

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