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Hayes and Garber - Cucurbit Breeding

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BREEDING OF VEGETABLES 245<br />

reactions are known indicate (McRostie, 1919; Burkholder,<br />

1918) that resistance to either the alpha or beta strain is inherited<br />

as a simple dominant, involving but a single factor difference.<br />

It seems, therefore, very easy to produce resistant varieties to<br />

both strains by crossing <strong>and</strong> selection <strong>and</strong> thus to combine desirable<br />

economic characters <strong>and</strong> anthracnose resistance.<br />

McRostie (1921) has recently published an interesting paper<br />

on further studies of disease resistance in common beans. The<br />

more extensive results obtained bear out the earlier views on<br />

the mode of inheritance of resistance to bean anthraxnose. The<br />

studies carried out show that bean mosaic susceptibility is inherited.<br />

In FI there was a partial dominance of susceptibility<br />

over resistance <strong>and</strong> in F z a segregation which indicated a two<br />

factor hypothesis.<br />

In crosses between susceptible <strong>and</strong> resistant<br />

varieties in relation to the dry root rot, caused by the fungus,<br />

Fusarium martii phaseoli Burk., there was a dominance in F\<br />

of susceptibility <strong>and</strong> a segregation in F% that appeared to be on<br />

a 9 :7 basis. In nearly all cases resistant F 2 plants bred true to<br />

this character in Fz. Results of this nature show the great<br />

practical importance of the application of Mendelian principles<br />

to breeding for disease resistance. It seems very likely that a<br />

large<br />

part of our serious plant diseases will be controlled eventually<br />

by the production of<br />

disease resistant varieties.<br />

TOMATO<br />

Classification Characters <strong>and</strong> Inheritance. The tomato belongs<br />

to the genus Lycopersicum of which there are several<br />

cultivated species. Tomatoes are classified on the basis of vine<br />

habit, either st<strong>and</strong>ard or dwarf, leaf type, period of maturity,<br />

size <strong>and</strong> color of fruits, <strong>and</strong> other characters. As a result of<br />

breeding experiments, many different combinations of characters<br />

have been made. Price <strong>and</strong> Drinkard (1908) were among the<br />

first investigators to report on the simple Mendelian behavior of<br />

certain tomato characters. Table LXIII, taken from similar<br />

ones compiled by Tschermak (1916) <strong>and</strong> Jones (1917), presents<br />

a brief summary of inheritance in the tomato.<br />

Fruit shape is dependent on several factors according to<br />

Crane (1915) <strong>and</strong> Groth (1912, 1915). Some of the foliage<br />

characters are also somewhat complicated in their inheritance<br />

(Groth, 1911). The inheritance of each of the other characters<br />

listed in the table is dependent on single factor differences.

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