Hayes and Garber - Cucurbit Breeding
Hayes and Garber - Cucurbit Breeding Hayes and Garber - Cucurbit Breeding
224 BREEDING CROP PLANTS 1. The heterozygous condition of most varieties. 2. The difficulties of obtaining crossed seed. The heterozygous condition need not be further emphasized. Conditions are much the same as in the fruit crops. The Difficulties of Obtaining Crossed Seed. The technic of making a cross is very simple. According to East (1908a), "The flowers close slightly about dusk and open in the morning between five and six o'clock. The pollen appears to be in the best condition for use on the second day of blooming." Stuart (1915) collects flowers to be used as the male parent in small sacks. After the pistil is removed from these flowers the anthers are tapped sharply with a pair of forceps, the pollen is collected FIG. 54. Emasculated and unemasculated potato blossoms. (After Stuart.) on the thumb nail and then applied to the pistil of the emasculated flower. The flowers are receptive two to four days after emasculation. East ( 1908a) stated the belief that the potato is usually self-fertilized. He also observed the fact that insects were seldom seen to visit the flower. Salaman (1910-11) believes it unnecessary to cover the flower before or after pollination. Stuart, however, used 1-lb. bags and found that if a certain amount of foliage was included in the bag the use of bags did not cause a lowering of the number of seeds set. An average of between one and two hundred seeds was obtained from each successful cross by Stuart. The chief difficulty is that many varieties do not bloom very
POTATO IMPROVEMENT 225 freely, although the general belief is that all varieties may bloom under certain conditions of environment. East (1908a) classified varieties as follows: "1. Varieties whose buds drop off without opening. " 2. Varieties in which a few flowers open, but which immediately fall. "3. Varieties whose flowers persist several days, but which rarely produce viable pollen. "4. Varieties which under most conditions always produce viable pollen," In 487 out of 721 varieties under observation the buds fell off before the flowers opened. Stuart, however, obtained a much higher percentage of varieties which produced flowers in which the blossoms opened before the buds fell. These results are given to emphasize the fact that conditions widely influence seed production. The lack of fertile or healthy pollen in many varieties prohibits their use as parents. The relation between the percentage of healthy pollen and fruit production was determined by East (1908a) for a considerable number of crosses (see Table LIV). TABLE LIV. RELATION BETWEEN PERCENTAGE OF VIABLE POLLEN AND FRUIT PRODUCTION Viability
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224 BREEDING CROP PLANTS<br />
1. The heterozygous condition of most varieties.<br />
2. The difficulties of obtaining crossed seed.<br />
The heterozygous condition need not be further emphasized.<br />
Conditions are much the same as in the fruit crops.<br />
The Difficulties of Obtaining Crossed Seed. The technic<br />
of making a cross is very simple. According to East (1908a),<br />
"The flowers close slightly about dusk <strong>and</strong> open in the morning<br />
between five <strong>and</strong> six o'clock. The pollen appears to be in the best<br />
condition for use on the second day of blooming." Stuart (1915)<br />
collects flowers to be used as the male parent in small sacks.<br />
After the pistil is removed from these flowers the anthers are<br />
tapped sharply with a pair of forceps, the pollen is collected<br />
FIG. 54. Emasculated <strong>and</strong> unemasculated potato blossoms. (After Stuart.)<br />
on the thumb nail <strong>and</strong> then applied to the pistil of the emasculated<br />
flower. The flowers are receptive two to four days after<br />
emasculation. East (<br />
1908a) stated the belief that the potato is<br />
usually self-fertilized. He also observed the fact that insects were<br />
seldom seen to visit the flower. Salaman (1910-11) believes<br />
it unnecessary to cover the flower before or after pollination.<br />
Stuart, however, used 1-lb. bags <strong>and</strong> found that if a certain<br />
amount of foliage was included in the bag the use of bags did<br />
not cause a lowering of the number of seeds set. An average of<br />
between one <strong>and</strong> two hundred seeds was obtained from each successful<br />
cross by Stuart.<br />
The chief difficulty is that many varieties do not bloom very