Hayes and Garber - Cucurbit Breeding
Hayes and Garber - Cucurbit Breeding Hayes and Garber - Cucurbit Breeding
222 BREEDING CROP PLANTS P. Segregating ratios were in accordance with the above factorial hypotheses. Wilson (1916) obtained only white tubers from selfed white-tubered varieties. Similar results have been obtained by other plant breeders which show that white is a recessive character. A certain amount of coloring in the young sprouts or shoots, stems, and sometimes in the leaf petioles was found associated with the presence of color in the tubers. With regard to flower color, three white-flowered varieties, selfed, produced only white flowers; and three out of four colored varieties, when selfed, produced both colored and white forms. Color is, therefore, dominant to its absence. Inheritance of this character may be explained by assuming the presence of a chromogen body and modifying factors. Heliotrope flowers are due to the chromogen body plus a reddening factor; purple flowers are produced by the addition of a purpling factor; white flowers may be due to the absence of one or more of these factors. Fruwirth (1912) found red tubers dominant over white, yellow flesh over white, and lilac-colored flowers over white. It was also found that different gradations of color were inherited. Nilsson (1912-13) found a complicated flower color inheritance. A variety with violet-blue flowers gave, on selfing, progeny with red, violet-blue, near-red, purple, dark and light blue, and white flowers. A variety with light blue flowers, on selfing, yielded progeny showing simple monohybrid segregation with white recessive. Evidence that several factors were operating in the inheritance of tuber flesh color was also obtained. Some of the varieties with yellow flesh (tubers) bred true when selfed, others segregated as dihybrids with white recessive. The inheritance of habit of growth was also studied. Plants may be upright, bushy, or procumbent. Bushy plants are heterozygous for habit of growth and many of them exhibit a distinct tendency to become procumbent. Homozygous forms of upright and sprawling plants may be isolated easily. maturity is used as a means of varietal classification. It Period of is probably inherited in the same manner as with other crops. Sterility of the anthers has been found to be a dominant character. At first Salaman believed that its inheritance was due to a single differential factor but later evidence indicated a more complex manner of transmission. Plants producing pale heliotrope flowers were found to be heterozygous for pollen sterility. MacDougal (1917) crossed the wild potato of Arizona, S.
fendleri, POTATO IMPROVEMENT 223 which grows at a high altitude and endures extremes of climate, with a domestic variety. The wild form produces small tubers. In the F 2 generation forms appeared which were identical with the wild parent together with many intermediate types. Most of the observed variations in cultivated varieties have occurred in the tubers, although the English ash-leaf varieties are examples of a variation in leaf shape (East, 19076). Production of New Forms. For the purpose of differentiating between two important phases of potato improvement, Stuart (1915) has referred to " selection" as the " isolation and asexual propagation of desirable strains or types" while " breeding" is used only for sexual reproduction. With certain crops, such as the potato, this terminology is distinctive. Such a restricted usage of the word "selection" seems undesirable from the plantbreeding standpoint. The same idea can be obtained by the use of "clonal selection" to refer to the asexual propagation of desirable strains or types. Systematic plant breeding with the idea of combining the desirable characteristics of two parental varieties can be carried out only after the breeder has familiarized himself with the characters of particular varieties and of their wild relatives. Thus, with the potato as with other crops the breeder should first determine the ideal toward which he will work. Parental varieties should than be selected because of some desirable characters. By recombination of the favorable characters of both parents, improvement may be obtained. Gilbert (1917) has listed certain characters of the potato which are universally desired. Some of these are: 1. High yield. 2. Good quality. 3. Disease-resisting capabilities. 4. Good keeping quality. 5. Good color of flesh and skin. 6. Skin of desirable texture. 7. Tubers of good shape. 8. Shallow eyes relatively few in number. 9. Upright, vigorous plants. 10. No tendency to make second growth. The desirability of most of these characters is self-evident. The chief difficulties in the way of developing a standardized method of attack arise from:
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fendleri,<br />
POTATO IMPROVEMENT 223<br />
which grows at a high altitude <strong>and</strong> endures extremes<br />
of climate, with a domestic variety. The wild form produces<br />
small tubers. In the F 2 generation forms appeared which were<br />
identical with the wild parent together with many intermediate<br />
types.<br />
Most of the observed variations in<br />
cultivated varieties have<br />
occurred in the tubers, although the English ash-leaf varieties<br />
are examples of a variation in leaf shape (East, 19076).<br />
Production of New Forms. For the purpose of differentiating<br />
between two important phases of potato improvement, Stuart<br />
(1915) has referred to " selection" as the " isolation <strong>and</strong> asexual<br />
propagation of desirable strains or types" while " breeding"<br />
is used only for sexual reproduction. With certain crops, such<br />
as the potato, this terminology is distinctive. Such a restricted<br />
usage of the word "selection" seems undesirable from the plantbreeding<br />
st<strong>and</strong>point. The same idea can be obtained by the use<br />
of "clonal selection" to refer to the asexual propagation of desirable<br />
strains or types.<br />
Systematic plant breeding with the idea of combining the<br />
desirable characteristics of two parental varieties can be carried<br />
out only after the breeder has familiarized himself with the<br />
characters of particular varieties <strong>and</strong> of their wild relatives.<br />
Thus, with the potato as with other crops the breeder should<br />
first determine the ideal toward which he will work. Parental<br />
varieties should than be selected because of some desirable<br />
characters. By recombination of the favorable characters of<br />
both parents, improvement may be obtained.<br />
Gilbert (1917) has<br />
listed certain characters of the potato which are universally desired.<br />
Some of these are:<br />
1. High yield.<br />
2. Good quality.<br />
3. Disease-resisting capabilities.<br />
4. Good keeping quality.<br />
5. Good color of flesh <strong>and</strong> skin.<br />
6. Skin of desirable texture.<br />
7. Tubers of good shape.<br />
8. Shallow eyes relatively few in number.<br />
9. Upright, vigorous plants.<br />
10. No tendency to make second growth.<br />
The desirability of most of these characters is self-evident.<br />
The chief difficulties in the way of developing a st<strong>and</strong>ardized<br />
method of attack arise from: