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Hayes and Garber - Cucurbit Breeding

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MAIZE BREEDING 183<br />

in the endosperm <strong>and</strong> a correspondingly large proportion of<br />

corneous starch. Some seeds may be entirely free from soft<br />

starch, but there is generally some soft starch surrounding, the<br />

germ. The group is characterized by the small size of its seeds<br />

<strong>and</strong> ears.<br />

Zea mays indentata, the dent corns. Sturtevant, Bulletin<br />

Torrey Botanical Club, 1894, page 329.<br />

The corneous starch in this group is located at the sides of the<br />

seed <strong>and</strong> the soft starch extends to the summit. The soft<br />

starch dries more rapidly than the corneous <strong>and</strong> this produces<br />

the shrinkage which causes the characteristic indentation of the<br />

seed.<br />

Zea mays amylacea, the soft or flour corns, Sturtevant, Bulletin<br />

Torrey Botanical Club, 1894, page 331.<br />

This group is recognized by an almost entire absence of corneous<br />

starch. There is no indentation in some varieties <strong>and</strong> only<br />

a slight one in others. The soft starch content characterizes<br />

this group.<br />

Zea mays saccharata, the sweet corns. Sturtevant, Bulletin<br />

Torrey Botanical Club, 1894, page 333.<br />

"A well-defined species group characterized by the translucent,<br />

horny appearance<br />

of the kernels <strong>and</strong> their more or less<br />

crinkled, wrinkled, or shriveled condition." East (1910d) presented<br />

evidence which shows that the sweet corns are dent, flint,<br />

or pop varieties which have not the ability to mature starch normally.<br />

The few starch grains produced are small, angular, <strong>and</strong><br />

imperfect.<br />

INHERITANCE OF CHARACTERS<br />

Endosperm Characters. The word xenia was first used by<br />

Focke (1881) to denote the effect which was apparently produced<br />

by the action of pollen upon the maternal tissue of the seed.<br />

The endosperm of maize was cited as a classical example of such<br />

an effect.<br />

After the discovery by Guignard (1899) <strong>and</strong> Nawaschin<br />

(1898) that the polar nuclei of the endosperm fuse with the<br />

second male nucleus of the pollen grain, De Vries (1899), Correns<br />

(1899), Webber (1900), <strong>and</strong> Guignard (1899, 1901) saw that this<br />

furnished an explanation of xenia in maize. From a consideration<br />

of inheritance of endosperm character the following law of<br />

xenia may be formulated :

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