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Hayes and Garber - Cucurbit Breeding

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CHAPTER XIV<br />

MAIZE BREEDING<br />

Maize was the most important bread crop of the American Indians<br />

<strong>and</strong> even today is the most important crop in the western<br />

hemisphere. The Indians brought the culture of maize to a<br />

high state of advancement <strong>and</strong> developed innumerable varieties.<br />

On the foundations made by the Indians modern corn-breeding<br />

has made marked advances, but perhaps no North American<br />

varieties are so notable as those developed by the Incas in Peru.<br />

Origin <strong>and</strong> Species. It is generally believed that Mexico is the<br />

original home of the maize plant, although there is no absolute<br />

proof of this (Harshberger, 1897). Zea mays L., belongs to the<br />

tribe Maydese of the order Graminese. All varieties of Indian<br />

corn are placed in the species mays. The nearest relative of<br />

maize is teosinte, Euchlcena mexicana Schrad. Teosinte <strong>and</strong><br />

maize cross readily <strong>and</strong> a natural hybrid between these cultivated<br />

grasses was described under the name Zea canina by Watson<br />

(Harshberger, 1904). A study of these crosses led Harshberger<br />

(1904, 1909) to make the hypothesis that maize originated from a<br />

hybrid between a sport of Euchlaena <strong>and</strong> normal teosinte. Montgomery<br />

(1906) reached the conclusion that maize <strong>and</strong> teosinte<br />

had a common progenitor. It was considered likely that the<br />

ancestral form of these cultivated grasses was a large muchbranched<br />

grass "each branch being terminated by a tassel-like<br />

structure bearing hermaphrodite flowers." As evolution progressed,<br />

the lateral branches of maize came to bear only pistillate<br />

flowers <strong>and</strong> the central branch staminate flowers. This theory<br />

is<br />

strengthened by the types of inflorescence which frequently<br />

appear in maize varieties. Often the central spike of the tassel of<br />

lateral branches bears seeds, while the side branches of the same<br />

tassel bear only staminate organs. All gradations appear<br />

between the normal ear of maize <strong>and</strong> the staminate tassel.<br />

It is<br />

not uncommon in self-fertilized maize races to obtain plants in<br />

which the tassel of the main branch bears both male <strong>and</strong> female<br />

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