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Hayes and Garber - Cucurbit Breeding

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166 BREEDING CROP PLANTS<br />

variability.<br />

A careful survey of experimental studies does not<br />

support this contention. The development of the shadegrown<br />

tobacco industry in the Connecticut Valley is of interest<br />

in this discussion. This shade method first originated in Florida<br />

in 1896 <strong>and</strong> was tried experimentally in Connecticut through<br />

cooperation of the Connecticut Experiment Station <strong>and</strong> officials<br />

of the Bureau of Soils. In 1900 one third of an acre was grown<br />

<strong>and</strong> the crop sold at an average price of 72 cents per pound.<br />

A considerable acreage was grown in 1901 <strong>and</strong> the crop sold<br />

at public auction at a much higher price per pound. Indiscriminate<br />

introduction of unselected seed from Florida was<br />

practiced <strong>and</strong> in 1902 over 700 acres were grown under shade in<br />

Connecticut. The result was a disastrous failure, owing to a<br />

lack of knowledge of methods of h<strong>and</strong>ling <strong>and</strong> to the use of unselected<br />

seed. By further study of h<strong>and</strong>ling <strong>and</strong> through careful<br />

selection in which artificially self-pollinated seed was saved,<br />

the industry was placed on a firm foundation. This latter work<br />

was carried on by the Bureau of Plant Industry (Stewart, 1908).<br />

A knowledge of Cuban methods shows that imported Cuban seed<br />

is a mixture of many types. Some experiments have shown that<br />

the breaking up alluded to is an expression of the different<br />

hereditary qualities of the parental seed plants. In 1912 Hasselbring<br />

grew a number of pure lines of tobacco in Michigan which<br />

he had formerly grown in Cuba. No evidence of breaking up of<br />

type was observed <strong>and</strong> whatever changes occurred in a pure line,<br />

owing to the new conditions, were uniformly exhibited in all<br />

plants of the pure line. Similar conclusions were reached from<br />

the immediate introduction of individual seed capsules of different<br />

tobacco plants from Cuba <strong>and</strong> their subsequent growth under<br />

shade in Connecticut (<strong>Hayes</strong>, 1914). Careful studies at Pusa,<br />

India, convinced the Howards (1910a) that new conditions<br />

did not cause a breaking up of type. They ascribed the apparent<br />

variability of new introductions to cross-fertilization, which<br />

was shown to occur frequently in tobacco.<br />

Although there have been some differences of opinion as to<br />

the cause of variability of new introductions, there is uniformity<br />

of belief regarding the methods of obtaining purity of type.<br />

Artificial self-pollination gives uniformity, <strong>and</strong> continued<br />

self-fertilization produces no harmful effects. This method was<br />

strongly recommended by officials of the United States Department<br />

of Agriculture (Shamel <strong>and</strong> Cobey, 1907) <strong>and</strong> by the different

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