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Hayes and Garber - Cucurbit Breeding

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COWPEAS, SOYBEANS, AND VELVET BEANS 147<br />

time of maturity are characters which have been widely used in<br />

varietal <strong>and</strong> group classifications.<br />

Little work has been done on the inheritance of characters in<br />

soybeans. Beans with green cotyledons may have green seedcoats,<br />

while beans with yellow cotyledons may have either green<br />

or yellow seed-coats. H. Terao 1<br />

(1918) of the Imperial Agricultural<br />

Experiment Station, Tokyo, Japan, has discovered that<br />

in a cross of green cotyledons, green seed-coats 9 X yellow<br />

cotyledons, yellow seed-coats cf the inheritance of the green<br />

seed-coat apparently was matroclinal ;<br />

likewise the inheritance of<br />

the character of the cotyledons. In the reciprocal cross the<br />

character of the cotyledons again proved matroclinal in inheritance<br />

but the seed-coat character segregated as a monohybrid<br />

with green dominant. In explanation of these facts it is assumed<br />

that the two kinds of chlorophyll concerned differ in that one remains<br />

green (G) <strong>and</strong> the other turns yellow (F). It is further<br />

assumed that the inheritance of these conditions in the cotyledons<br />

is through the cytoplasm or chromatophores <strong>and</strong> not through<br />

the nucleus. In the case of color of seed-coat a Mendelian factor<br />

pair is involved. When H is present it prevents the chlorophyll<br />

(F) in the seed-coat from changing to yellow.<br />

is absent the small letter h is used.<br />

When this factor<br />

Table XXXIII. taken from Terao illustrates four possible<br />

combinations. (G) <strong>and</strong> (F) are transmitted only through the<br />

cytoplasm of the egg cell.<br />

TABLE XXXIII. INHERITAN<br />

Parents<br />

Cotyledons.<br />

-<br />

IN i<br />

CR<<br />

(G)HH<br />

cTppn

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