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Hayes and Garber - Cucurbit Breeding

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134 BREEDING CROP PLANTS<br />

A Wheat Cross Made at Svalof. The highest yielding winter<br />

wheat grown at the Svalof Station, reported by Newman (1912),<br />

was a cross, Extra Squarehead II, No. 0290. This wheat is<br />

one of the offspring of Old Extra Squarehead X Grenadier II.<br />

It combines the winter-hardiness <strong>and</strong> rust resistance of the former<br />

with the stiff straw <strong>and</strong> high yield of the latter. As an average<br />

of four years' trial at Svalof <strong>and</strong> Alnarp, this wheat has yielded<br />

18 per cent, more than Old Extra Squarehead <strong>and</strong> 8 per cent,<br />

more than Grenadier II, which was next. No variety of winter<br />

wheat has proved so generally popular among the farmers of<br />

southern Sweden as Extra Squarehead II.<br />

It may be of interest<br />

to point out that preceding the cross, hundreds of selections out<br />

of Grenadier II were examined in search of a pure line with the<br />

combination of rust resistance <strong>and</strong> high yield.<br />

Wheat <strong>Breeding</strong> at University Farm, Cambridge, Engl<strong>and</strong>.<br />

Most of the wheat varieties grown in Engl<strong>and</strong> are very susceptible<br />

to yellow rust (Pucdnia glumarum). Biffen (1917) set himself<br />

the task of breeding a high-yielding, resistant form. He crossed<br />

American Club, which is very resistant to this parasite, with<br />

several susceptible varieties in order to study the mode of inheritance<br />

<strong>and</strong> develop a st<strong>and</strong>ard technic of operations. In all<br />

crosses the F% generations showed monohybrid segregation with<br />

resistance behaving as the recessive. The resistant individuals<br />

were rather clear-cut, although they sometimes exhibited uredinia.<br />

The susceptible plants showed a wide range of variation. No<br />

recognizable morphological character has been found correlated<br />

with resistance.<br />

The constancy of resistance in wheats of hybrid origin has also<br />

been studied by Biffen. For the purpose he used a resistant<br />

strain produced from a cross between American Club, a resistant<br />

variety, <strong>and</strong> Michigan Bronze, which is one of the forms most<br />

susceptible to yellow rust. During eight years of observation<br />

the hybrid variety proved just as resistant as the American Club.<br />

A resistant variety of Russian origin, found among some<br />

Gurka wheats, which was not adapted to local conditions, was<br />

crossed with Square Head's Master, the variety most commonly<br />

grown in Engl<strong>and</strong>. Among the resistant offspring is one that<br />

gives considerable promise. Comparative trials of this wheat<br />

(Little Joss) over a period of seven years show it to yield about 4<br />

bu. per acre more than the best of the English <strong>and</strong> French wheats.<br />

The explanation for this would seem to be that Little Joss in-

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