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Hayes and Garber - Cucurbit Breeding

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96 BREEDING CROP PLANTS<br />

susceptibility in F 3 ,<br />

while resistant F 2 plants were of two kinds:<br />

(1) those which produced only resistant progeny <strong>and</strong> (2) those<br />

which segregated, both resistant <strong>and</strong> susceptible plants being<br />

obtained.<br />

Size Characters. Nilsson-Ehle (1908) made numerous studies<br />

of inheritance of size characters. In a cross between two<br />

saliva varieties which differ in height, transgressive segregation<br />

occurred in F 2 . Forms were selected <strong>and</strong> the studies continued<br />

FIG. 22. Culms of resistant <strong>and</strong> susceptible varieties of oats. From left to<br />

right: Victory, susceptible to stem rust; a susceptible F% plant of Victory X<br />

White Russian; a resistant Fz plant of Victory X White Russian; resistant White<br />

Russian.<br />

through F 4 <strong>and</strong> F b .<br />

Segregation was of a complex nature.<br />

Transgressive segregation also occurred in crosses involving leaf<br />

breadth, kernel size, <strong>and</strong> number of florets to the spikelet. The<br />

results were explained on the multiple factor hypothesis, but the<br />

actual factors involved could not easily be determined. Maturity<br />

may be considered under this heading, for it behaves in a similar<br />

manner. From crossing early <strong>and</strong> later maturing oats', Caporn<br />

(1918) obtained intermediate maturity in FI <strong>and</strong> segregation<br />

in F*. The author suggests that three factors will quite satis-

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