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Hayes and Garber - Cucurbit Breeding

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CLASSIFICATION AND INHERITANCE OF SMALL GRAINS 95<br />

<strong>and</strong> in F 2 the articulated basal types could easily be determined.<br />

These occurred in a close approximation of the ratio of 1 articulated<br />

base to 3 of the intermediate <strong>and</strong> saliva types (Fraser,<br />

1919).<br />

Open versus Side Panicle. Nilsson (1901) has used panicle<br />

types <strong>and</strong> seed colors as a chief means of classification. The<br />

distinction between the side <strong>and</strong> the open panicle is easily made,<br />

but the various transitional open panicled forms are not easily<br />

used in differentiation. Nilsson-Ehle (1908) has explained<br />

crosses between an open-panicled <strong>and</strong> a side-panicled variety on<br />

the basis of two main factor differences. Either factor when<br />

homozygous or heterozygous produces open panicles. When<br />

both factors are homozygous a variety with an open panicle <strong>and</strong><br />

drooping branches is obtained. When the factors are absent a<br />

side panicle results. From crossing two open-panicled forms,<br />

9-side forms were obtained out of a total of 112 plants.<br />

These<br />

side-panicled plants bred true while of the 103 open-panicled<br />

plants, 24 again segregated giving both open- <strong>and</strong> side-panicled<br />

forms. The parental varieties have panicles with erect branches<br />

while a part of the open-panicled segregates have drooping<br />

branches.<br />

Resistance to Rust. Parker (1918) studied varietal resistance<br />

of oats to stem rust, Puccinia graminis avence Erikss. <strong>and</strong> Henn.<br />

<strong>and</strong> to crown rust, Puccinia lolii avence McAlpine. Crown rust<br />

is a serious disease in the South while stem rust is more common<br />

in the North. Several varieties of the red oat group of A.<br />

sterilis including Burt, proved resistant to crown rust, while<br />

certain side oat strains of A. saliva orienlalis belonging to the<br />

White Russian group proved resistant to stem rust.<br />

Studies, of the inheritance of resistance to crown rust under<br />

greenhouse conditions, of crosses of Burt with Sixty Day, A.<br />

saliva, showed segregation in F 2 .<br />

Susceptible <strong>and</strong> resistant<br />

plants, as well as various intermediates, were obtained (Parker,<br />

1920).<br />

A study of the inheritance of resistance to stem rust has<br />

been made at the Minnesota Station (<strong>Garber</strong>, 1921). FI, F 2 ,<br />

<strong>and</strong> F z crosses of resistant White Russian with two susceptible<br />

varieties of A. saliva, Victory <strong>and</strong> Minota, have been grown.<br />

The preliminary results show that for these crosses resistance is a<br />

dominant character, the ratio in F 2 of resistant <strong>and</strong> susceptible<br />

plants approximating 3:1. Susceptible F 2 plants bred true to

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