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Hayes and Garber - Cucurbit Breeding

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CLASSIFICATION AND INHERITANCE IN WHEAT 87<br />

disease causes an annual decrease of 15 per cent, in the yield of<br />

winter wheat in the States of Washington, Oregon, <strong>and</strong> Idaho.<br />

The method used in studying bunt resistance was to blacken<br />

the seed with smut spores just before planting <strong>and</strong> then sow the<br />

strains <strong>and</strong> crosses in rows in such a way that each plant could<br />

be individually examined.<br />

Studies of inheritance were made in crosses of Turkey X Florence<br />

<strong>and</strong> Turkey X Hybrid 128. Hybrid 128 is a prolific,<br />

winter-hardy, stiff-strawed wheat of much commercial value<br />

but it is<br />

very susceptible to bunt. Turkey does not yield as well<br />

as Hybrid 128, has weak straw, <strong>and</strong> shatters considerably. It is<br />

highly resistant to bunt. Florence is an Australian spring wheat<br />

which is highly resistant to bunt.<br />

The results presented b}^ Gaines show very clearly that resistance<br />

to bunt is an inherited character.<br />

However, several factors<br />

are necessary to explain the sort of segregation obtained. The F 2<br />

of the cross between Florence <strong>and</strong> Turkey showed transgressive<br />

segregation. In Fa, 171 families were grown from individual F 2<br />

plants. Of these, 72 were immune while 50 families produced over<br />

80 per cent, of bunt. The Turkey <strong>and</strong> Florence parents under<br />

the same conditions produced an average of 4.6 per cent, of<br />

infected plants. This shows that the factors for resistance in the<br />

Florence <strong>and</strong> Turkey varieties are not identical.<br />

In the cross between Turkey <strong>and</strong> Hybrid 128 no segregates<br />

were obtained with a higher degree of resistance than the Turkey<br />

parent. It was found possible to produce resistant strains of<br />

any morphological type desired.<br />

Inheritance of Other Characters. Nilsson-Ehle (IQllc, 1912)<br />

has shown that winter-hardiness is inherited in much the same<br />

manner as other characters. Segregation occurs in F% <strong>and</strong> types<br />

can be produced in later generations which are homozygous for<br />

different degrees of winter-hardiness. Crosses made (<strong>Hayes</strong> <strong>and</strong><br />

<strong>Garber</strong>, 1919) in 1902 between hardy Odessa winter wheat <strong>and</strong><br />

Turkey varieties were bred for several years by continuous selection<br />

methods. Odessa is a late maturing variety <strong>and</strong> does not<br />

give a high yield in Minnesota. Turkey is a desirable winter<br />

wheat in many sections but it lacks hardiness under Minnesota<br />

conditions. Two early maturing wheats, Minhardi <strong>and</strong> Minturki,<br />

have been produced from the cross between Turkey <strong>and</strong><br />

Odessa. These new varieties excel in winter-hardiness <strong>and</strong><br />

yield.

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