Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...
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Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...

Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ... Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...

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19.11.2013 Views

78 ADVANCES IN PALAEOICHTHYOLOGY

ACTA UNIVERSITATIS LATVIENSIS, 2004, Vol. 679, pp. 79-98 A new tetrapod Jakubsonia livnensis from the Early Famennian (Devonian) of Russia and palaeoecological remarks on the Late Devonian tetrapod habitats OLEG A. LEBEDEV Oleg A. Lebedev, Palaeontological Institute of the Russian Academy of Sciences, 123 Profsoyuznaya St., Moscow 117997, Russia; olebed@paleo.ru Skeletal remains of the earliest tetrapod Jakubsonia livnensis gen. et sp. nov. are described from the Lower Famennian, Upper Devonian of the Gornostayevka quarry (SW of the Livny Town, Oryol Region, Russia). A part of the skull roof, incomplete angular, the right cleithrum with scapulocoracoid and partial femur are described. The new genus demonstrates characters in the structure of the interorbital region and the cleithrum suggesting similarity to the Ventastega Ahlberg, Lukševics and Lebedev, 1994. An analysis of early tetrapod sites indicates that these animals dwelled within a wide range of aquatic environments. The antiarch genus Remigolepis played an important role in Famennian tetrapod communities. The presence of more than one tetrapod in the communities indicates many more diverse tetrapod trophic adaptations than previously considered. Key words: tetrapods, Russia, Oryol region, Devonian, Famennian, palaeoecology, Jakubsonia. Introduction Studies of Late Devonian tetrapods became especially intensive during the last decade of the 20th century. Apart from Ichthyostega Säve-Söderbergh, 1932 and Acanthostega Jarvik, 1952, new taxa from most continents of the world, of both Laurussian and Gondwanan origin, have recently been described. Most of these tetrapods (Metaxygnathus Campbell and Bell, 1977, from Australia, Tulerpeton Lebedev, 1984 from Russia, Hynerpeton Daeschler, Shubin, Thomson and Amaral, 1994 and Densignathus Daeschler, 2000 from Pennsylvania (U.S.A.), Ventastega Ahlberg, Lukševics and Lebedev, 1994 from Latvia, Sinostega Zhu, Ahlberg, Zhao and Jia, 2002 from China and an ichthyostegid tetrapod from Belgium) (Campbell and Bell 1977; Lebedev 1984; Lebedev and Clack 1993; Daeschler et al. 1994; Lebedev and Coates 1996; Ahlberg, Lukševics and Lebedev 1994; Daeschler 2000; Zhu et al. 2002; Lukševics, Ahlberg and Clack 2003; Clement et al. 2004) originate from the upper and middle Famennian deposits. Only Elginerpeton pancheni Ahlberg, 1995 is known from the upper Frasnian of Scotland (Ahlberg 1995, 1998). For that reason, each new find of tetrapod material in deposits older than middle Famennian make a contribution of special interest to our knowledge in this field of vertebrate palaeontology. The first finds of vertebrates from the rich fossil vertebrate locality Gornostayevka quarry (sometimes previously called the Livny quarry) situated to the SW of the Livny

78 ADVANCES IN PALAEOICHTHYOLOGY

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