Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...

Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ... Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...

19.11.2013 Views

34 ADVANCES IN PALAEOICHTHYOLOGY Genus Perscheia gen. nov. Etymology. In honor of Per Schei, the geologist on the 1898-1902 Fram Expedition, who collected the only previously known psammosteids from the Canadian Arctic. Perscheia pulla sp. nov. (Figs. 6-10) Etymology. From the Latin, pullus, black, referring to the black color of the specimens. Diagnosis. Dorsal plate thick and with surface ornamentation of well developed elongated blisters and irregular radial ridges. Histological structure of spongy aspidin reinforced by pleromin towards the outer surface. Ventral plate thick and with long posterior median notch. Material. Six pieces of plate from the dorsal and ventral armor: CMN-NUFV 101-CMN- NUFV106. Holotype. Posterior part of the dorsal plate (CMN-NUFV101). Locality. Lower part of the Nordstrand Point Formation, southern Ellesmere Island, inland from Okse Bay. N77 o 06.163’ W87 o 09.064’. Field site number: NV2K 11. Description. The material consists of five large pieces of dorsal or ventral plate and one small fragment that has been sectioned to show the histology. The holotype, CMN–NUFV101 (Figs. 6A, 6B, 7A), is part of a plate showing a natural edge and three broken edges. It is convex on the outer and concave on the inner surfaces, roughly square (with about one quarter missing), and 180 mm in length and 210 mm in width. The thickness varies from 5 mm at the natural edge to 18.5 mm on the opposite edge. The external surface is finely pitted and ridged due to the presence of the spongy aspidin layer at the surface. It is covered by irregular blisters or excrescences on the more medial part of the plate; this ornament is replaced about 100 mm from the natural margin of the plate by low, round-topped, irregular, radial ridges. The change of ornament occurs at what appears to be a growth line delineated by a groove that runs parallel to the natural edge. A similar growth line is present about 45 mm from the plate edge. The ridges are generally about 3-5 mm wide, the blisters are mostly about 5 mm x 8 mm becoming larger and more pronounced towards the central part of the plate. Two sensory canals run across the plate at right angles to the natural edge and parallel to the ridges (Fig. 7A). The more complete of these runs the entire length of the specimen. It forms a shallow groove 0.5-0.75 mm wide and has four short lateral branches in the zone of blisters, but only one in the ridged zone. A longer lateral branch runs parallel to the growth line that delineates the change in ornament. At the broken edge of the plate the longitudinal canal connects to a sensory canal running at right angles to it along the edge of the specimen. From this canal three additional short branches project. The longitudinal canals are probably the medial dorsal canals, the canal at right angles forming a dorsal transverse commissure.

D.K. Elliott, E. Mark-Kurik, E.B. Daeschler. A revision of Obruchevia 35 The inner face of the specimen is smooth and shiny, and at least three growth lines, delineated by either shallow grooves or thickening of the bone, run parallel to the natural edge (Fig. 6B). The surface is formed of a dense laminar basal layer of aspidin that is up to 2 mm thick and tends to flake away from the spongy overlying layers. The surface shows small foramina related to narrow, shallow, radial grooves and large smooth blisters that are elongated radially. This specimen is assumed to be part of the median dorsal plate, probably the median posterior part based on the presence of the medial canals, the orientation of the ornament, and the shape of the natural margin. CMN–NUFV102 (Figs. 6C, 7B) is a roughly pentagonal, concavo-convex fragment of plate in which all the edges are broken. Its maximum dimensions are 134 mm x 112 mm and the thickness varies from 15 mm to 10 mm. The outer surface of the plate is ornamented with large elongated blisters, apparently radially oriented. Several of these are ring shaped with a depressed center. Four sections of the sensory canal system can be seen, forming two parallel pairs almost at right angles to each other (Fig. 7B). One pair consists of a short section within the plate and a section along the right-hand edge. They run at an angle to the trend of the ridges, the canal on the plate edge does so almost at right angles to the ridges. The second pair consists of a canal that runs along the longest edge, almost at right angles to the first pair and a short section on the opposite edge. The longest section of canal is almost parallel to the trend of the ornament suggesting that it is part of the medial dorsal canal; the short section parallel to it may be part of the lateral dorsal canal, while the other two sections of canals are probably transverse commissures. The internal surface is concave, smooth, and shiny with a thick, dense, laminar, basal layer and no obvious growth lines. CMN–NUFV103 (Figs. 6D, 7D) is an irregular fragment with maximum dimensions of 96 mm and 161 mm. It varies in thickness from 12 mm to 7 mm. The external surface is covered by large blisters and short ridges arranged roughly parallel to the long axis of the plate fragment. At about the midpoint of the specimen a sensory canal runs almost at right angles to the trend of the ridges (Fig. 7D). Six short lateral branches separate from the main canal and project between the surface ridges. Towards one end a short section of canal can be see running along one edge and parallel to the radial ornament. This canal is probably one of the transverse commissures while the longer section is part of one of the lateral or medial dorsal canals. The internal surface is weathered and the laminar basal layer is missing, exposing the spongy aspidin above it. CMN-NUFV104 (Figs. 6E-G, 7C) is an irregular and thick piece of plate, 85 mm x 155 mm and up to 28 mm thick. It has broken margins on all edges except for one small section 55 mm long on one margin where there is a natural rounded edge to a shallow embayment. The curved margin adjacent to the embayment is probably close to the plate margin as it thins rapidly. The outer surface is concavo-convex with a suggestion of growth lines parallel to the curved margin (Fig. 6F). A single sensory canal is present running across the middle of the fragment for two-thirds of its width and then branching at right angles (Fig. 7C). One branch runs parallel to the embayment to the broken margin in one direction and for about 50 mm parallel to the plate margin in the other direction before petering out. Presumably this part of the sensory canal is the lateral dorsal canal. Although damaged the surface shows the same pattern of coarse blisters present on the other plates.

D.K. Elliott, E. Mark-Kurik, E.B. Daeschler. A revision of Obruchevia<br />

35<br />

The inner face of the specimen is smooth <strong>and</strong> shiny, <strong>and</strong> at least three growth lines,<br />

delineated by either shallow grooves or thickening of the bone, r<strong>un</strong> parallel to the<br />

natural edge (Fig. 6B). The surface is formed of a dense laminar basal layer of aspidin<br />

that is up to 2 mm thick <strong>and</strong> tends to flake away from the spongy overlying layers. The<br />

surface shows small foramina related to narrow, shallow, radial grooves <strong>and</strong> large smooth<br />

blisters that are elongated radially. This specimen is assumed to be part of the median<br />

dorsal plate, probably the median posterior part based on the presence of the medial<br />

canals, the orientation of the ornament, <strong>and</strong> the shape of the natural margin.<br />

CMN–NUFV102 (Figs. 6C, 7B) is a roughly pentagonal, concavo-convex fragment<br />

of plate in which all the edges are broken. Its maximum dimensions are 134 mm x 112 mm<br />

<strong>and</strong> the thickness varies from 15 mm to 10 mm. The outer surface of the plate is<br />

ornamented with large elongated blisters, apparently radially oriented. Several of these<br />

are ring shaped with a depressed center. Four sections of the sensory canal system can<br />

be seen, forming two parallel pairs almost at right angles to each other (Fig. 7B). One<br />

pair consists of a short section within the plate <strong>and</strong> a section along the right-h<strong>and</strong> edge.<br />

They r<strong>un</strong> at an angle to the trend of the ridges, the canal on the plate edge does so<br />

almost at right angles to the ridges. The second pair consists of a canal that r<strong>un</strong>s along<br />

the longest edge, almost at right angles to the first pair <strong>and</strong> a short section on the<br />

opposite edge. The longest section of canal is almost parallel to the trend of the ornament<br />

suggesting that it is part of the medial dorsal canal; the short section parallel to it<br />

may be part of the lateral dorsal canal, while the other two sections of canals are<br />

probably transverse commissures. The internal surface is concave, smooth, <strong>and</strong> shiny<br />

with a thick, dense, laminar, basal layer <strong>and</strong> no obvious growth lines.<br />

CMN–NUFV103 (Figs. 6D, 7D) is an irregular fragment with maximum dimensions of<br />

96 mm <strong>and</strong> 161 mm. It varies in thickness from 12 mm to 7 mm. The external surface is<br />

covered by large blisters <strong>and</strong> short ridges arranged roughly parallel to the long axis of<br />

the plate fragment. At about the midpoint of the specimen a sensory canal r<strong>un</strong>s almost<br />

at right angles to the trend of the ridges (Fig. 7D). Six short lateral branches separate<br />

from the main canal <strong>and</strong> project between the surface ridges. Towards one end a short<br />

section of canal can be see r<strong>un</strong>ning along one edge <strong>and</strong> parallel to the radial ornament.<br />

This canal is probably one of the transverse commissures while the longer section is<br />

part of one of the lateral or medial dorsal canals. The internal surface is weathered <strong>and</strong><br />

the laminar basal layer is missing, exposing the spongy aspidin above it.<br />

CMN-NUFV104 (Figs. 6E-G, 7C) is an irregular <strong>and</strong> thick piece of plate, 85 mm x 155<br />

mm <strong>and</strong> up to 28 mm thick. It has broken margins on all edges except for one small<br />

section 55 mm long on one margin where there is a natural ro<strong>un</strong>ded edge to a shallow<br />

embayment. The curved margin adjacent to the embayment is probably close to the<br />

plate margin as it thins rapidly. The outer surface is concavo-convex with a suggestion<br />

of growth lines parallel to the curved margin (Fig. 6F). A single sensory canal is present<br />

r<strong>un</strong>ning across the middle of the fragment for two-thirds of its width <strong>and</strong> then branching<br />

at right angles (Fig. 7C). One branch r<strong>un</strong>s parallel to the embayment to the broken<br />

margin in one direction <strong>and</strong> for about 50 mm parallel to the plate margin in the other<br />

direction before petering out. Presumably this part of the sensory canal is the lateral<br />

dorsal canal. Although damaged the surface shows the same pattern of coarse blisters<br />

present on the other plates.

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