Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...
Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...
Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...
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D.K. Elliott, E. Mark-Kurik, E.B. Daeschler. A revision of Obruchevia<br />
27<br />
Halstead Tarlo (1964, p. 20; 1965, p. 149) explained in detail the problems concerning<br />
the generic name. He published the family name, Obrucheviidae Tarlo, 1964, generic<br />
name Obruchevia Whitley, 1940, <strong>and</strong> species name Obruchevia heckeri (Obruchev,<br />
1936) as the valid ones (Halstead Tarlo 1965); however, he did not refer to Obruchev’s<br />
paper of 1940. Obruchev <strong>and</strong> Mark-Kurik (1965, p. 19, 78) used the name Aspidosteus,<br />
<strong>and</strong> mentioned Obruchev’s paper of 1940. They did not, however, describe the genus<br />
Aspidosteus as the monograph concerned only the representatives of the family<br />
Psammosteidae Traquair.<br />
Obruchev could not <strong>and</strong> did not consider the name Obruchevia Whitley, 1940 as a<br />
nomen nudum as Whitley correctly followed the Article 13 of the International Code of<br />
Zoological Nomenclature (1964), referring to Obruchev’s paper of 1936. Obruchev (1964)<br />
did not ignore Whitley’s (1940) corrections of fish names in other cases, either. For<br />
example, he recognized Whitley as the author of the generic name Cyrtaspidichthys for<br />
the genus Cyrtaspis Bryant, 1932. Whitley anticipated White <strong>and</strong> Moy-Thomas (1940),<br />
who published a new name, Eucryptaspis, for Bryant’s genus only one month later (in<br />
J<strong>un</strong>e 1940).<br />
In many papers published after 1941 on Devonian biostratigraphy (e.g., Blieck et<br />
al. 1988) <strong>and</strong> fish paleontology of the NW of the East European platform, including<br />
Obruchev’s own papers (e.g., that of 1967), the name Aspidosteus (not Obruchevia)<br />
was commonly used. It was, however, not the case with Halstead Tarlo who regularly<br />
used the name Obruchevia in his publications (Halstead Tarlo 1963, p. 3; 1967b, p.<br />
1233; Halstead 1969, p. 22; 1974, p. 61 etc.) but not yet in Halstead Tarlo 1962 (p. 261).<br />
Therefore, in relation to the priority problem, the stability or <strong>un</strong>iversality criteria<br />
cannot be applied (see Article 23 in the International Code of Zoological Nomenclature,<br />
1999).<br />
There seem to be two possibilities: (1), Obruchev considered his description of<br />
Aspidophorus (later called Aspidosteus or Obruchevia) in 1936 not valid as publication<br />
was in a popular science journal (Priroda); or (2), he published the name Aspidosteus in<br />
some paper before May of 1940, i.e. before the correction was made by Whitley (however<br />
we have not fo<strong>un</strong>d such a paper). Halstead Tarlo (1964, p.124; 1965, p. 159) referenced<br />
two of Obruchev´s papers in press (Obruchev, D., 1965a, “On the branchial<br />
plates of Aspidosteus,” <strong>and</strong> Obruchev, D., 1965b, “Pycnosteus nathorsti n. sp. from the<br />
Middle Devonian of Spitsbergen”) but <strong>un</strong>fort<strong>un</strong>ately, neither was ever published.<br />
It is concluded here that as Obruchev (1964, p. 75 in the Russian edition) clearly<br />
indicated that the genus Aspidosteus was established by him in 1941, the priority belongs<br />
to the name Obruchevia Whitley, 1940, <strong>and</strong> the name Aspidosteus Obruchev,<br />
1941 is its j<strong>un</strong>ior synonym.<br />
Localities. Obruchev (1964) mentioned that Aspidosteus was known from the Novgorod<br />
Region, Russia, <strong>and</strong> from Latvia. According to Lukševičs (2001) Aspidosteus occurs in<br />
Latvia in the Pamūšis Regional Stage, whereas in Lyarskaya <strong>and</strong> Lukševičs (1992) it was<br />
mentioned as occurring in two <strong>un</strong>its: the Katleši <strong>and</strong> Ogre (= Pamūšis) formations.<br />
Lukševičs (pers. comm. 2003) is of the opinion that the occurrences of Aspidosteus from<br />
Latvia were after all erroneously reported.<br />
The main distribution area of Obruchevia is the Novgorod Region, NW Russia<br />
(Fig. 2, upper left). Six Obruchevia (Aspidosteus) localities have been reported on the<br />
River Lovat´ about 30 km upstream from the railway bridge of the connection between