Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...
Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ... Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...
112 ADVANCES IN PALAEOICHTHYOLOGY they are loosely attached to the head shield. Only three of the shields are complete consisting of unpaired Nu, Pp and Prm plates, as well as paired Pn, Pmg and La plates. Two head shields lack Pmg from one side, and four more shields are without Pmg plates from both sides. Three head shields have lost the anterior most portion, and the Prm plate is not present. In some cases (specimens LDM 81/283, 81/726) it seems that the head shield has been disarticulated after burial, since plates are displaced and situated at a distance of 1 to 3 mm from each other. One of the best preserved portions of the trunk shield is specimen LDM 81/434, consisting of articulated right ADL, AVL, Cd1 and Cv1. Besides this specimen, there are 18 examples of articulated AVL plates and proximal segments of pectoral fin. Specimen No. LDM 81/716, for example, shows articulated AVL plates on both sides with connected proximal segments. Another seven specimens are articulated proximal segments of pectoral fin alone. Only two specimens, LDM 81/58-60 and 81/83, have a partially preserved dorsal wall of the trunk shield. Three specimens (LDM 81/41, 81/81 and 81/82) constitute an incomplete ventral wall of the trunk shield. By counting articulated or partly articulated portions of shields of Bothriolepis one may infer that the disintegration of placoderm fish skeletons took place quite close to the place of burial, and that the final stages of disintegration continued even after transportation was finished. Sarcopterygian and tetrapod remains are represented with some completely disarticulated bones of the shoulder girdle and skull, as well as rare scales or sometimes teeth, but more or less complete lower jaws have often been found, e.g. three partial lower jaws of Holoptychius cf. nobilissimus, five jaws of Cryptolepis grossi, and 14 complete or partially complete jaws of Ventastega curonica. There are two almost complete specimens representing tetrapod cheeks, and one skull roof of Ventastega, which was found at a distance of some 45 cm from the left side cheek. Judging from the size and shape of broken edges, both skull and cheek belong to the same individual. Skeletal element representation. The sorting of remains according to their shape, size and weight has been observed. Among the remains, medium-sized (4-6 cm long) bones prevail, smaller remains such as scales of Cryptolepis grossi are disproportionally rare, but larger ones such as lower jaws of Ventastega curonica (about 21 cm long) are considerably more frequently found than the other skeletal elements of the animal (Fig. 7 A). Voorhies assigned skeletal elements of some modern mammals to one of three categories in accordance with their ease of fluvial transport: Group I is the most readily transported and Group III the most resistant to transport (Behrensmeyer 1975). It is impossible to conduct the Voorhies Group analysis of the Devonian vertebrate assemblages for two reasons. Firstly, skeletons of different taxa in a mixed taxonomic assemblage yield different numbers of elements for each Voorhies Group, and sometimes homologous elements from even phylogenetically close taxa fall into different Voorhies Groups (Behrensmeyer 1975). Secondly, placoderms and ancient tetrapods have no direct analogues with homological skeletal elements among modern taxa. Therefore the abilities of different skeletal elements of vertebrates from the Pavāri oryctocoenosis to be transported and sorted by fluvial processes can only be evaluated. A most satisfactory analysis is possible only examining representation of Bothriolepis ciecere, from which almost all bones are represented in the assemblage. Ventastega curonica
E. Lukševičs, I. Zupiņš. Sedimentology, fauna, and taphonomy of Pav āri 113 and Cryptolepis grossi could also give satisfactory results since they are represented by many skeletal elements. It is, however, impossible to use the other taxa due to the insufficient number of specimens. Disarticulated bones of the trunk and head shield of Bothriolepis ciecere are represented in different numbers. Strongly convex dorsolateral and ventrolateral bones, particularly posterior ventral lateral plates from the left side (PVL) and mixilateral plates from the right side (MxL), dominate among the trunk shield bones (Tab. 3). Possibly these bones, along with the less convex anterior median dorsal (AMD), could belong to the Voorhies Group III, being the most resistant to transport. The number of the anterior ventro-lateral (AVL) and anterior dorso-lateral (ADL) plates is smaller, and the almost flat posterior median dorsal (PMD) and flat median dorsal (MV) are the rarest among the trunk shield bones, representing the Voorhies Group II, which is moderately readily transported. Besides that, the distribution of paired convex bones shows assymetry. The number of the right-sided MxL plates more than three times exceeds the number of the left-sided MxL, but the situation is opposite in the case of PVL plates. The number of the left-sided PVL two times exceeds the number of the right-sided PVL. The differences between various Voorhies groups and number of bones from the right and left side of the trunk suggest quite an important hydraulic sorting of skeletal elements of placoderms. Bones of the head shield are also represented in different numbers, always being more rare than the trunk shield plates. Thick and relatively large lateral (La) and nuchal (Nu, n = 17) plates, as well as complete head shields, are found comparably often. Still, articulated head shields, which supposedly could represent the Voorhies Group III, are disproportionally rare. Separate premedian (Prm, n = 2) and other small and thin bones are very rare. Such very thin and small bones as inferognathals and submarginals, as well as bones of sklerotic ossifications, were not found. Possibly the lightest bones were readily transported soon after death. Table 3. Number of some skeletal elements of placoderm fish Bothriolepis ciecere * Unpaired bones and shield
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E. Lukševičs, I. Zupiņš. Sedimentology, fa<strong>un</strong>a, <strong>and</strong> taphonomy of Pav āri<br />
113<br />
<strong>and</strong> Cryptolepis grossi could also give satisfactory results since they are represented<br />
by many skeletal elements. It is, however, impossible to use the other taxa due to the<br />
insufficient number of specimens.<br />
Disarticulated bones of the tr<strong>un</strong>k <strong>and</strong> head shield of Bothriolepis ciecere are<br />
represented in different numbers. Strongly convex dorsolateral <strong>and</strong> ventrolateral bones,<br />
particularly posterior ventral lateral plates from the left side (PVL) <strong>and</strong> mixilateral<br />
plates from the right side (MxL), dominate among the tr<strong>un</strong>k shield bones (Tab. 3).<br />
Possibly these bones, along with the less convex anterior median dorsal (AMD), could<br />
belong to the Voorhies Group III, being the most resistant to transport. The number of<br />
the anterior ventro-lateral (AVL) <strong>and</strong> anterior dorso-lateral (ADL) plates is smaller,<br />
<strong>and</strong> the almost flat posterior median dorsal (PMD) <strong>and</strong> flat median dorsal (MV) are the<br />
rarest among the tr<strong>un</strong>k shield bones, representing the Voorhies Group II, which is<br />
moderately readily transported. Besides that, the distribution of paired convex bones<br />
shows assymetry. The number of the right-sided MxL plates more than three times<br />
exceeds the number of the left-sided MxL, but the situation is opposite in the case of<br />
PVL plates. The number of the left-sided PVL two times exceeds the number of the<br />
right-sided PVL. The differences between various Voorhies groups <strong>and</strong> number of bones<br />
from the right <strong>and</strong> left side of the tr<strong>un</strong>k suggest quite an important hydraulic sorting of<br />
skeletal elements of placoderms. Bones of the head shield are also represented in different<br />
numbers, always being more rare than the tr<strong>un</strong>k shield plates. Thick <strong>and</strong> relatively large<br />
lateral (La) <strong>and</strong> nuchal (Nu, n = 17) plates, as well as complete head shields, are fo<strong>un</strong>d<br />
comparably often. Still, articulated head shields, which supposedly could represent the<br />
Voorhies Group III, are disproportionally rare. Separate premedian (Prm, n = 2) <strong>and</strong><br />
other small <strong>and</strong> thin bones are very rare. Such very thin <strong>and</strong> small bones as inferognathals<br />
<strong>and</strong> submarginals, as well as bones of sklerotic ossifications, were not fo<strong>un</strong>d. Possibly<br />
the lightest bones were readily transported soon after death.<br />
Table 3. Number of some skeletal elements of placoderm fish Bothriolepis ciecere<br />
* Unpaired bones <strong>and</strong> shield