66 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 253 Fig. 34. Distribution <strong>of</strong> Liophryne rhododactyla (circles) and L. similis (triangle) in Papua New Guinea. Shaded area is greater than 2400 m elevation. downward behind the ear rather than continuing straight onto the flank. DESCRIPTION OF HOLOTYPE: The holotype specimen is a mature female (2-mm ova), with the following measurements and pro- portions (those <strong>of</strong> the single paratype given in parentheses): SVL 37.5 (37.0), HW 15.0 (14.5), TL 21.1 (21.0), EY 4.0 (4.1), EN 2.8 (2.8), IN 4.9 (4.3), TY 3.2 (2.8), HD 8.3 (8.0), FT 19.9 (18.4), disc <strong>of</strong> third finger 1.1
2000 ZWEIFEL: PARTITION OF <strong>SPHENOPHRYNE</strong> 67 Fig. 35. Holotype <strong>of</strong> Liophryne rubra, UPNG 9290, SVL 37.5 mm. (0.09), penultimate phalanx 0.06 (0.05), disc <strong>of</strong> fourth toe 1.7 (1.4), penultimate phalanx 0.8 (0.08), TY 3.2 (2.8); HW/SVL 0.400 (0.392), TL/SVL 0.563 (0.568), EY/SVL 0.107 (0.111), EN/SVL 0.075 (0.076), IN/ SVL 0.131 (0.116), EN/IN 0.571 (0.651), HD/SVL 0.221 (0.216), FT/SVL 0.528 (0.497), FD/SVL 0.029 (0.024), TD/SVL 0.045 (0.038). Head narrower than the rather robust body, snout bluntly pointed seen from above, rounded and slightly projecting in pr<strong>of</strong>ile; nostrils lateral, barely visible from above, slightly closer to eye than to tip <strong>of</strong> snout; canthus rostralis distinct, slightly rounded; loreal region steep, nearly flat. Eyes moderate in size, corneal outline visible from beneath, interorbital span about 1.5 width <strong>of</strong> eyelid. Tympanum relatively large and smooth. Relative lengths <strong>of</strong> fingers 4 3 2 1, first relatively long, reaching to base <strong>of</strong> disc <strong>of</strong> second, all with discs broader than penultimate phalanges; subarticular elevations moderately prominent, rounded. Relative lengths <strong>of</strong> toes 4 3 5 2 1, first short, reaching base <strong>of</strong> subarticular elevation <strong>of</strong> second, all with discs broader than penultimate phalanges, disc <strong>of</strong> fourth toe about Fig. 36. Holotype <strong>of</strong> Liophryne rubra, UPNG 9290. 1.5 width <strong>of</strong> that <strong>of</strong> third finger; subarticular elevations moderately prominent, rounded; inner metatarsal elevation low, rounded. A straight fold <strong>of</strong> skin passing from posterior corner <strong>of</strong> eye diagonally on flank where it fades out; skin otherwise smooth with no dorsal tubercles or scapular folds. There appear to be small maxillary teeth. Shortly after being preserved, the specimen was ‘‘bright chestnut red’’ in color (J. Menzies, personal commun.); the dorsal surfaces (head, body, legs, and feet) have since faded to pale yellowish tan, unmarked except for a white vertebral hairline. A dark streak passes from nostril to eye and resumes at the posterior corner <strong>of</strong> the eye to include much <strong>of</strong> the ear before terminating (fig. 36). The streak is sharply defined at the canthus rostralis, less so along its lower edge; the upper lip is pale. The ventral surfaces are pale and unmarked from chin to hind legs. There are a few dark marks on front and rear surfaces <strong>of</strong> the front legs, an irregular longitudinal dark streak on the anterior side <strong>of</strong> each thigh, and a poorly defined dark streak on the posterior side. The palms and soles have some dark pigment but are not wholly dark. VARIATION IN TYPE SERIES: The paratype resembles the holotype in all pertinent characters, most notably the relatively short eye– naris span, absence <strong>of</strong> convergent scapular folds, and color pattern. Body proportions are summarized in table 6. REFERRED SPECIMEN: A specimen <strong>of</strong> Liophryne 9 from the Baiyer River region some 9 SAMA R23853, collected by Thomas Burton Jan. 3, 1982, at Trauna Ridge, 1750 m, 13 km E Baiyer River Wildlife Sanctuary, Western Highlands Prov., Papua New Guinea.